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Status Accounts for Selected Threatened Indian Mammals


Order ARTIODACTYLA

Family Bovidae

GAUR
Status: VULNERABLE
Bos gaurus (H. Smith, 1827)CITES: Appendix I

The Gaur, or Indian Bison, inhabits evergreen and deciduous forested hills and associated grassy clearings up to 1,800 m, eastwards from India, Nepal and Bhutan to Burma, Thailand, Laos, Cambodia, Vietnam, southern China and the Malay Peninsula. A large (650 - 1,000 kg) and where undisturbed mainly diurnal animal, it lives in herds of from 2 - 40 individuals containing only one mature bull. Other adult males may live singly or in bachelor groups. Primarily a grazer, the Gaur prefers green grass but in its absence will take coarse dry grass, forbs, and the leaves of some trees and creepers. In eastern India and Burma a domesticated form of the Gaur, the Mithan, is widely used both as a draught animal and for meat production. The wild population probably numbered at least 10,000 in 1983 (Thornback 1983). It was however declining drastically through over-hunting; intolerance of habitat destruction and disturbance by people; and exposure to the diseases of domestic cattle (e.g. rinderpest, foot-and-mouth and anthrax). Outside Protected Areas the Gaur is now confined to remote and isolated forest regions. The species breeds in captivity, and is protected by law in most countries of its range. Adequate law enforcement in all Protected Areas in which the Gaur persists is required.

Sources:

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

Thornback, J. (1983). Wild Cattle, Bison and Buffaloes: Their Status and Potential Value. Publ. by WCMC, Cambridge, U.K.

WILD YAK
Status: ENDANGERED
Bos mutus Linnaeus, 1766CITES: Appendix I ( = Bos grunniens )

The Wild Yak inhabits alpine tundra and ice desert in the Tibetan plateau and adjacent highlands at altitudes from 4,270 to 6,100 m, occurring in China, India and possibly Bhutan, Nepal and Afghanistan. Males may weigh up to 1,000 kg, females only one-third as much. The long shaggy coat reaches almost to the ground, and in spite of the Yak's awkward appearance it is an agile climber. It stays in high areas with permanent snow during the few summer months, and descends to lower elevations during the rest of the year. The diet consists of grasses, herbs and lichens. Females and young live in large herds - once reported to contain thousands of individuals - while adult males live alone or in groups of up to 12, only joining the females during the rut. The Domesticated Yak is an integral part of the economy of the people of the area, used as a draught animal and for milk, meat and wool production. Although legally protected in China and India, Wild Yak have declined drastically in the past few decades - primarily through uncontrolled hunting by local tribes - and the species is now confined to scattered and remote areas of its former range. In 1983 the wild population was estimated to be in the low hundreds compared to a population of Domesticated Yaks numbering millions (Thornback 1983). Measures considered necessary to conserve the Wild Yak include surveys to locate viable populations; legal protection throughout its range; the establishment of further Protected Areas; and conservation education programmes.

Sources:

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

Thornback, J. (1983). Wild Cattle, Bison and Buffaloes: Their Status and Potential Value. Publ. by WCMC, Cambridge, U.K.

WILD ASIATIC WATER BUFFALO
Status: ENDANGERED
Bubalus arnee (Linnaeus, 1758)CITES: Appendix III (= Bos bubalus)

The Wild Asiatic Water Buffalo naturally occurs only in India (mainly in Assam and Madhya Pradesh) and Nepal. Wild populations found elsewhere in Asia, Australasia and Brazil are the descendants of either translocated or feral individuals, since some 130 million Domestic Water Buffalo are employed in agriculture throughout the world. The Wild Asiatic Water Buffalo weighs 700 - 1,200 kg and inhabits wet grasslands, swamps, densely vegetated river valleys and lake margins, and sometimes drier savannah with scattered trees, particularly in the south of its range. It is chiefly a grazer, feeding in the morning and evenings and lying up in dense cover or submerged in wallows during the heat of the day. Females and young usually associate in small groups, often accompanied by a single adult bull, but led by an old cow. Other males live solitarily or in bachelor herds. Once common, the species' decline has been caused by loss of its riverine habitat to cultivation and human settlement; interbreeding with Domestic Water Buffalo; and competition for forage with, and vulnerability to disease (particularly rinderpest) transmitted by, domestic stock. By the 1980s the total population was probably only 1,000 - 1,500 (Thornback 1983) scattered mainly in various Protected Areas. Due to interbreeding with free-ranging Domestic Water Buffalo only some 100 of these in the Bastaar region of Madhya Pradesh could be considered "pure" stock (Kane, 1989). Various proposed hydroelectric and irrigation projects threaten some of the last known haunts of the species. The Asiatic Wild Water Buffalo will soon become extinct unless measures are taken to protect its habitat and segregate populations from livestock to prevent interbreeding and disease transmission. It is protected by law in both India and Nepal, but none are kept in captivity.

Sources:

Kane, R. (1989). The Wild Buffalo. Asian Wild Cattle Specialist Group Newsletter, No. 2. pp 4 - 5.

Thornback, J. (1983). Wild Cattle, Bison and Buffaloes: Their Status and Potential Value. Publ. by WCMC, Cambridge, U.K.

MARKHOR
Status: ENDANGERED
Capra falconeri (Wagner, 1839) CITES: Appendix II

HB 140-168cm; SH 65-100cm; W 32-100kg; H 82-143cm (MacDonald 1985).

The Markhor was once widespread in rugged mountain ranges within the north-western great Himalayan Chain of Afghanistan, Pakistan and India, its range also extending in a southerly arc down through the mountain ranges of Waziristan and Baluchistan in Pakistan. The largest species of goat, with adult males weighing up to 100kg, it is a striking animal since both sexes bear long sharp-keeled horns twisted into a tight or open spiral, and in addition males have a long neck mane, pantaloons and strong markings. It has been eliminated over most of its range by over-hunting as a trophy animal, particularly within the last 25 years, and also suffers from competition with domestic stock. It now survives mainly in small populations in isolated areas. Originally seven subspecies were distinguished but taxonomic revision has reduced this to three (Schaller and Khan 1975): the Flare-horned Markhor C. f. falconeri from eastern Afghanistan, northern Pakistan and north-western India; the Tadzhik Markhor C. f. heptneri of Uzbekistan and Tadzhikistan in southern USSR and the adjacent Badakhshan area of northern Afghanistan; and the Straight-horned Markhor C. f. megaceros of the border area between Afghanistan and Pakistan, from approximately the Khyber Pass to Quetta. This last subspecies is classed as Endangered and is dealt with separately below. The Markhor occurs in forest steppe and Artemesia steppe country at altitudes of from 700 - 4,000 m, with access to cliffs (which form its refuge from predators) and suitable terrain below 2,200 m where temperatures are moderate in winter. In 1976 the total population of Markhor was estimated as 8,000, although precise estimates are difficult because of the physical and political inaccessibility of many parts of its range. The breakdown was as follows: 4500 C. f. falconeri in Afghanistan and Pakistan plus 250-300 in India: less than 2,000 C. f. megaceros, and 1,000 C. f. heptneri in USSR plus 125-150 in Afghanistan (IUCN 1976). It is believed that since 1976 most populations have declined, with the exception of that in Hazarganji-Chiltan National Park which has increased under governmental protection, and one small population north of Quetta, which has increased after being accorded protection by the local community (Thornback, 1989). The species is totally protected by law in India and the USSR, and partially protected in some states of Pakistan. It occurs in several Protected Areas, including Chitral Gol Reserve in Pakistan and the Dashti-Dhum Sanctuary in Tadzhikistan - which harbours 70% of the population of the USSR - but there is an urgent need to increase the number of Reserves and improve levels of protection. Once populations are re-established, controlled trophy hunting could be implemented to generate revenue. The Markhor breeds readily in captivity and is a popular exhibit. The species is listed on CITES Appendix II, with 3 subspecies C. f. chialtaensis, C. f. jerdoni, and C. f. megaceros (see below) listed on Appendix I.

Sources:

IUCN, (1976). Markhor Red Data Sheet. IUCN, Gland, Switzerland, and Cambridge, UK.

Macdonald, D.W. (1984). The Encyclopaedia of Mammals. Unwin Hyman, London, UK. 895 pp.

Thornback, J. (1989). Markhor Capra falconeri in Pakistan. Unpublished report to N.C.C. 3 pp + Map.

NILGIRI TAHR
Status: VULNERABLE
Hemitragus hylocrius Ogilby, 1838 CITES: Not listed

HB 90-140cm; SH 61-106cm; T 9-12cm; H 45cm; W 50-100 kg (Nowak 1991).

The Nilgiri Tahr is a goat-like animal with a short coat and short, laterally-flattened, curved horns. Males are black with a silver saddle and a short bristly mane, while females are greyish brown with white bellies. It is active intermittently from dawn to dusk and is primarily a grazer, living in herds of from 6 - 104 animals, with average group sizes of 9 for all-female groups and 27 for mixed herds (Davidar 1978, Schaller 1977). Mating takes place throughout the year but there is a birth peak in winter. Wild tahr rarely give birth to twins. A single offspring is born after a gestation period of 180 - 242 days, and females can give birth twice in one year. The species was once abundant in grass-woodland mosaic habitat in rugged hills and mountain slopes at altitudes from 1200 - 1600 m in the southern Indian states of Kerala and Tamil Nadu. It is now limited to some 17 populations in the Nilgiri, Anamalai, Palani and Highwavy Hills, the Eravikulam area of the High Range, and possibly a few other areas in the western Ghats. In 1986 total numbers were estimated at 2,000 - 2,200 (Rice, 1986) relatively unchanged since 1978 (Davidar 1978). The Nilgiri Tahr has been protected by law in India since 1972 and the two largest populations (comprising approximately 50% of the remaining animals) occur in Nilgiri Hills and Eravikulam National Parks. In 1987 numbers in the latter were estimated at 550 (Rice, 1988). Most other populations are small (20 - 100) and isolated, and deserve more attention since all are subject to poaching by shooting and snaring, and many are also threatened by grazing competition with domestic stock, hydroelectric schemes, and habitat loss to agriculture and plantations of eucalyptus and wattle. Ecological studies are needed to form the basis of management plans since sustainable harvesting and / or licensed sport hunting of a restored population could be a valuable source of protein and income for local people. The Nilgiri Tahr breeds well in captivity.

Sources:

Davidar, E. R. C. (1978). Distribution and status of the Nilgiri Tahr (Hemitragus hylocrius) 1975-1978. J. Bombay Nat. Hist. Soc. 75: 815-844.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

Rice, C. G. (1986). Conservation of Tahr. Caprinae News. No. 1 (1): 7-9.

Rice, C. G. (1988). Reproductive biology of Nilgiri Tahr Hemitragus hylocrius (Mammalia: Bovidae). J. Zool. Lond. 214, 269-284.

Schaller, G. B. (1977). Mountain Monarchs. Wild sheep and goats of the Himalaya. Univ. Chicago Press, Chicago, USA.

Family Suidae

PYGMY HOG
Status: ENDANGERED
Sus salvanius (Hodgson, 1847)CITES: Appendix I

At 6 -10 kg the smallest member of the family Suidae, the Pygmy Hog was once found in the southern Himalayan foothill belt in southern Nepal, northern Bangladesh, Sikkim, Bhutan and adjacent parts of northeast India. Considered extinct by the 1960s, it was rediscovered in 1971 in north-western Assam in India, to which its distribution now seems confined. The population is extremely small - in 1978 total numbers were estimated at only 100 - 150 (Oliver in press). The Pygmy Hog inhabits tall-grass upland savanna or "thatchland", lives in small family groups of 5 - 20 individuals, and feeds mainly on roots, tubers, insects and small mammals. It is under threat from permanent habitat loss to human settlement, cultivation, and afforestation; temporary habitat loss due to the harvesting of grasses to provide local thatching materials, and the officially-sanctioned burning of areas of grassland to improve grass yield for thatch and grazing - both activities which temporarily displace the Hog into less favourable regions; and hunting for meat. Although the species itself is fully protected under Indian law, enforcement is difficult. In addition its habitat is only safeguarded in two Protected Areas: Barnadi Wildlife Sanctuary (2,621 ha) the site of the Pygmy Hog's original "reappearance", and Manas Wildlife Sanctuary which probably harbours the only viable wild population. A plan to construct a hydroelectric dam and flood control scheme in Manas was defeated in 1986 after the intervention of the Prime Minister of India, but the Sanctuary was then occupied by armed political extremists of the All Bodo Students Union in late 1988 / early 1989 (Oliver, 1993). The current status of the Pygmy Hog in Manas is therefore not known since access is impossible, but it is assumed to be critical. The establishment of new Protected Areas, improved protection of wild populations, and better management of captive populations are necessary. The Pygmy Hog breeds well in captivity. A three-point Action Plan proposed by the IUCN Pigs and Peccaries Specialist Group is to be implemented in Barnadi, rather than in Manas as originally suggested, and a field study of the species' behavioural ecology and a captive breeding programme are planned for 1991.

Source:

Oliver, W. (1993). Pigs, Peccaries and Hippos. Status Survey and Conservation Action Plan. IUCN, Gland, Switzerland.

Order CARNIVORA

Family Canidae

GREY WOLF
Status: VULNERABLE
Canis lupus Linnaeus, 1758CITES: Appendices I & II

The Grey Wolf was once the world's most widely distributed terrestrial mammal, occurring throughout the Palaearctic region above 20oN, and ranging southwards into northern Mexico, Saudi Arabia and peninsular India. Populations in the USA, Canada, former USSR and Poland are large and relatively stable - for example the Canadian population was estimated at 45,000 - 65,000 in 1991 (Anon. 1991) - but elsewhere numbers are decreasing due to a combination of habitat destruction by human settlement and agriculture; hybridisation with domestic dogs; and persecution as a livestock predator, competitor for game species (particularly deer), and for fear of attacks on humans. It is now extinct in many parts of its original European range (e.g. U.K., France and Germany) and only some 5,750 - 6,250 are left in west and central Europe (Ginsberg & Macdonald 1990). At 12 - 80 kg the Grey Wolf is the largest canid species. It occupies a variety of habitats from Arctic tundra through forest to desert. Home range size varies from 30 to 1,000 km2 depending on the habitat occupied. Living in large family packs, its diet consists mainly of large ungulates which are hunted co-operatively although smaller prey items, carrion and garbage may also be taken (Mech 1970). The legal status of the Grey Wolf varies greatly from region to region. In some areas it is completely protected, whilst in others it is the subject of concerted eradication efforts by bounties, poisoning and government control programmes. Although substantial populations exist in Protected Areas in some parts of its range, conservation measures necessary in Europe include full legal protection; introduction of livestock-compensation schemes for herders; limiting of development in protected areas; reintroduction of prey species; control of feral dog populations and establishment of captive breeding programmes. The Grey Wolf breeds well in captivity. Populations in Bhutan, India, Nepal and Pakistan are listed on Appendix I of CITES, all others are on Appendix II.

Sources:

Anon. (1991). World Wildlife Fund - Canada proposes carnivore conservation strategy. Wolf! Vol. IX, No. 1: 29-31.

Ginsberg, J. R. and Macdonald, D. W. (1990). Foxes, Wolves, Jackals and Dogs. An Action Plan for the Conservation of Canids. IUCN, Gland, Switzerland.

Mech, D. L. (1970). The Wolf. New York: Natural History Press.

DHOLE
Status: VULNERABLE
Cuon alpinus (Pallas, 1811)CITES: Appendix II

The Dhole, Red Dog or Asiatic Wild Dog is a fairly large (10 - 20 kg) pack-living canid found in thick scrub jungles and dense forests up to 3000 m throughout mainland southeast Asia from India to eastern China, and on the islands of Java and Sumatra. Dholes prey mainly on medium-sized ungulates e.g. chital deer, wild boar and wild sheep, but rodents, reptiles, insects and berries are also taken. The population is in steep decline in most parts of India, the Soviet Union and the Himalayas due to disease (virulent canine distemper and rabies) exacerbated by contact with domestic dogs; deforestation; depletion of natural prey outside Reserves; and poisoning by herders. Fortunately reasonably large sub-populations exist in Protected Areas in India, where they have been studied by Johnsingh (1982, 1985). In the long-term, populations outside Protected Areas are unlikely to survive. Recommended conservation steps include assessments of the status of the species in Burma (which possibly harbours the largest remaining wild population) the former USSR and China, and the banning of the use of poisons in Grey Wolf control programmes in the former USSR. Dholes are legally protected in both India and the Soviet Union.

Sources:

Ginsberg, J. R. and Macdonald, D. W. (1990). Foxes, Wolves, Jackals and Dogs. An Action Plan for the Conservation of Canids. IUCN, Gland, Switzerland.

Johnsingh, A. J. T. (1982). Reproductive and social behaviour of the dhole (Cuon alpinus). Journal of Zoology, (London) 198:443-463.

Johnsingh, A. J. T. (1985). Distribution and status of the dhole Cuon alpinus Pallas, 1811 in South Asia. Mammalia 49:203-208.

BENGAL FOX
Status: INDETERMINATE
Vulpes bengalensis Shaw, 1800CITES: Appendix III

The Bengal fox inhabits open country, thorny scrub and semi-desert up to 1,500 m in India, Pakistan and Nepal. Weighing 1.8 - 3.2 kg, it is believed to form long-term monogamous pairs, and feeds opportunistically on insects, fruits, ground-nesting birds and smaller mammals. The species has been eliminated from most of the Indian plains by killing for sport and for the alleged medicinal properties of its flesh, coupled with the loss of open habitat to agriculture and human settlement. Currently no significant populations exist in Protected Areas in India, but the establishment of reserves for the Great Indian Bustard Ardeotis nigriceps should improve the situation. An education campaign aimed at curbing sport killing is necessary, together with a survey of the distribution and status of the species in Nepal and Pakistan.

Source:

Ginsberg, J. R. and Macdonald, D. W. (1990). Foxes, Wolves, Jackals and Dogs. An Action Plan for the Conservation of Canids. IUCN, Gland, Switzerland.

Family Felidae

ASIATIC CHEETAH
Status: ENDANGERED
Acinonyx jubatus venaticus (Griffith, 1821)CITES: Appendix I

The Asiatic subspecies of the Cheetah is classed as Endangered by IUCN. Originally distributed from the Middle East to India and extending northwards into the former USSR, it has disappeared from most of its range. It is believed to survive in desert steppes and low hills in the plateau provinces of Iran and possibly also occurs in adjacent parts of Pakistan, Afghanistan and Turkmenistan. Although long thought to be extinct in India one sighting was reported in November 1990 from Simlipal Wildlife Sanctuary (Anon, 1991). Like its African relative it has suffered from persecution as a predator of livestock, and the decimation of its natural prey species - particularly gazelles - by motorized sport hunting. The Asiatic Cheetah is totally protected by law in Iran and the former USSR, and until 1977 populations in Protected Areas in Iran were reported to be stable or increasing. There were then estimated to be 200 remaining in the wild. In the last decade however, the population has decreased severely. The present number, status and distribution of Cheetah in Iran is not known, and should be clarified.

Sources:

Anon. (1991). Cheetah sighting in India. Oryx 25: 69.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

ASIATIC GOLDEN CAT
Status: INSUFFICIENTLY KNOWN
Catopuma (= Felis) temmincki (Vigors & Horsfield, 1827)CITES: Appendix I

At 12 - 15 kg one of the larger members of the genus Felis, the Asiatic Golden Cat is widely distributed from Tibet, Nepal and India to southern China, Indochina, Thailand, the Malay peninsula and Sumatra. It mainly inhabits dry deciduous forest and tropical rainforest, though it is occasionally found in more open habitats. It is mainly terrestrial, often hunts in pairs, and preys on hares, small deer, birds, lizards and other small mammals. Nowhere common in any part of its range, the species is declining in India and Indonesia due to habitat destruction and intolerance of human disturbance. It is protected by law in Malaysia and in most Indian states. A survey of the status, distribution and ecology of the Asiatic Golden Cat is required to clarify its situation.

Sources:

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

IUCN (1978). Asiatic Golden Cat Red Data Sheet. IUCN, Gland, Switzerland and Cambridge, U.K.

CLOUDED LEOPARD
Status: VULNERABLE
Neofelis nebulosa (Griffith, 1821)CITES: Appendix I

The Clouded Leopard inhabits jungle and forests up to 2500 m from northeastern India and Nepal to southern China, Taiwan, the Malay peninsula, Sumatra and Borneo. A medium-sized cat weighing 16 - 23 kg, it is believed to live in pairs which may hunt together. It preys on birds, monkeys, pigs, cattle, young buffalo, goats, deer and porcupines. Its numbers have declined drastically in the last 10 - 20 years due to habitat loss to agriculture and overhunting for its pelt. Although protected by law in most countries, illicit trade in skins continues. In Taiwan there is a growing market for pelts smuggled from mainland China, since a Clouded Leopard jacket is a traditional symbol of prestige. The species breeds well in captivity.

Sources:

IUCN (1978). Clouded Leopard Red Data Sheet. IUCN, Gland, Switzerland and Cambridge, U.K.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

ASIATIC LION
Status: ENDANGERED
Panthera leo persica (Mayer, 1826)CITES: Appendix I

Once widely distributed from northern Greece eastwards through Syria, Iraq and Iran to central India, the Asiatic Lion was exterminated over most of its range with the advent of modern firearms in the 19th Century. The last remaining population occurs in the 1412 km2 Gir Forest Sanctuary of Gujarat State in India, where it survived under the patronage of the Nawab of Junagadh, and is now strictly protected under Indian law. One of 7 subspecies of the Lion (Panthera leo) - 2 of which are already extinct - the Asiatic Lion weighs 150 - 250 kg and inhabits teak forest, scrub and grassland. It lives in social groups or prides and cooperatively hunts large mammals e.g. nilgai, sambar, chital, and wild pig. At one time the Gir Forest suffered heavily from overgrazing by domestic cattle, reducing natural ungulate populations and forcing the lions to turn to livestock predation. Cattle have now been effectively excluded and the lion population has increased from 180 in 1974 to 280 in 1990 (Anon, 1990). Plans have been proposed to create a second lion reserve, both to relieve the pressure on the Gir Sanctuary and to lessen the risks of a catastrophe wiping out a single population.

Sources:

Anon. (1990). Asiatic Lion population up. p12 in Cat News 13.

IUCN (1978). Asiatic Lion Red Data Sheet. IUCN, Gland, Switzerland, and Cambridge, U.K.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

TIGER
Status: ENDANGERED
Panthera tigris (Linnaeus, 1758)CITES: Appendix I

HB 140-280cm; T 60-95cm; W 65-384kg (Nowak, 1991).

Within the past century the Tiger occurred from eastern Turkey and the Caspian Sea across Central Asia to the sea of Okhotsk, and south through the Indian sub-continent to South East Asia, Sumatra, Java and Bali. It is adapted to a wide range of habitats, from tropical rain forest, evergreen forest and mangrove swamp to grassland, savanna and rocky country. The largest species of cat, weighing from 65 - 384 kg, the Tiger is usually solitary and preys mainly on large mammals such as pigs, deer, antelope, buffalo, gaur and on occasion, man. Fond of water, Tigers can swim long distances. Male territories overlap those of several females. The Tiger now persists only in isolated pockets of its former range. Its decline has been caused by habitat loss, mainly to agriculture, logging, and malaria eradication programmes; overhunting (both legal and illegal) for sport, for skins, and for carcasses which are highly sought after for use in Asian traditional medicines; elimination of natural prey; and deliberate eradication programmes and persecution for fear of attacks on humans and domestic livestock - which have tended to increase as numbers of wild prey have dwindled. Incidental capture in traps set for wild pig and deer, and the fragmentation and isolation of populations resulting in poor breeding success have become more important threats as the numbers of Tigers have decreased. Of the original 8 subspecies of Tiger, 3 (the Caspian Tiger P. t. virgata, the Bali Tiger P. t. balica and the Javan Tiger P. t. sondaica) are believed to be Extinct and the remaining 5 are considered Endangered. The total number of Tigers in the wild in 1990 was only 6,500 - 9,000 (Jackson 1990) and probably only 2 subspecies now have viable wild populations. The Indian Tiger P. t. tigris increased in India from 1,800 in 1972 to 4,300 in 1990 under the government's "Project Tiger" programme to establish Tiger Reserves (Anon, 1990) and in 1990 the total population numbered 4,500 - 6,000; while that of the Indochinese Tiger P. t. corbetti was estimated at 1,100 - 1,500 (Jackson in litt.). In contrast the South China Tiger P. t. amoyensis is in immediate danger of extinction, having declined from an estimated 4,000 in 1949, to 150 - 200 in 1981 - 1982, and being further reduced to 30 - 50 in 1990 (Koehler, 1991). The populations of the Amur or Siberian Tiger P. t. altaica (estimated at 380 - 480 in 1990) and the Sumatran Tiger P. t. sumatrae (estimated at 500 - 1,000 in 1990) are fragmented and severely threatened (Jackson 1990). The Tiger is totally protected by law in most countries of its range except Burma, and occurs in many Protected Areas (e.g. Corbett Tiger Reserve, India). It breeds well in captivity and a Tiger Studbook is produced each year. In the 1990 edition the captive population was recorded as 1085 individuals belonging to all 5 subspecies (Anon. 1991). A Global Tiger Master Plan is also in preparation to conserve the species in the wild with support from captive breeding and reintroduction programmes. Other measures necessary include the effective enforcement of existing laws by increasing levels of patrolling in Protected Areas to suppress poaching both of the Tiger and its natural prey; a clampdown on illegal trade in skins and tiger products; prevention of encroachment on existing Reserves and the continued establishment of new ones; and a greater emphasis on protection outside Reserves. In some regions public education and information campaigns, and research on Tigers and their prey may also be necessary (Koehler 1991). All populations of the Tiger are now listed on Appendix I of CITES.

Sources:

Anon. (1990). Results of Tiger Census in India. Cat News 13: 3.

Anon. (1991). International Tiger Studbook 1990. Cat News 15: 6.

Jackson, P. (1990). In litt. to WCMC.

Koehler, G. M. (1991). Survey of Remaining Wild Population of South China Tigers. WWF Project 4512 / China. Final Project Report. WWF, Gland.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

MARBLED CAT
Status: INSUFFICIENTLY KNOWN
Pardofelis (= Felis) marmorata Martin, 1837CITES: Appendix I

The Marbled Cat inhabits thick jungle from Nepal, India and northern Burma to Thailand, Indochina, the Malay peninsula, Sumatra and Borneo. A small (2 - 6 kg) shy, nocturnal species, it is thought to prey on birds, squirrels, rats and other small mammals, lizards, snakes, frogs and small monkeys. It has always been regarded as rare, but this may reflect the inaccessibility of its habitat and its secretive nature. Protected by law in most parts of its range, the Marbled Cat is threatened by habitat destruction and disturbance, since it deserts forests that are even moderately disturbed. A survey of the distribution, status and ecology of the species is required to clarify its situation.

Sources:

IUCN (1978). Marbled Cat Red Data Sheet. IUCN, Gland, Switzerland and Cambridge, U.K.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

RUSTY-SPOTTED CAT
Status: INSUFFICIENTLY KNOWN
Prionailurus (= Felis) rubiginosa (I. Geoffroy, 1831)CITES: Appendices I & II

The Rusty-Spotted Cat is a small (1 - 2 kg) solitary species found in scrub, dry grassland, and open country in India, and humid mountain forest in Sri Lanka. It is nocturnal and preys on birds and small mammals. Nothing is known about its status or potential threats to its survival. It has been recorded from the Gir Forest Sanctuary in India. The Indian population is on Appendix I of CITES, the Sri Lankan on Appendix II.

Source:

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

SNOW LEOPARD
Status: ENDANGERED
Uncia (= Panthera) uncia (Schreber, 1775)CITES: Appendix I

The Snow Leopard (or Ounce) is found in the high altitude mountain ranges of Central Asia from Afghanistan, Pakistan and the former USSR in the west through northern India, Nepal and Bhutan to Mongolia and China. In summer it inhabits the arid alpine meadow region between the tree-line and permanent snow at elevations from 2,700 to 6,000 m, while in winter it may follow its prey down into the forests below 1,800 m. At 25 - 75 kg the smallest member of the genus Panthera, the Snow Leopard is solitary, crepuscular, and preys on mountain ungulates such as markhor, ibex, bharal and tahr, domestic stock, and smaller mammals such as hares and marmots. It is believed to be in decline in most regions due to uncontrolled hunting (for its fur and other trophies, as vermin because it preys on livestock, and because a large cat represents a challenge to the hunting instinct of many hill people); increased use of alpine pastures by people and their livestock; and a reduction in natural prey. The world population was recently estimated as 5 - 6,000 (Jackson 1987) with some 2,000 of these in China (Schaller 1988) and 500 in India (Fox et al. 1991). Although the Snow Leopard is legally protected in all countries of its range, enforcement is difficult in the remote terrain it inhabits, and therefore a stricter ban on the bazaar trade in skins is necessary. It is present in several Protected Areas (e.g. Hemis National Park, India) but more are needed to secure refuges both for the cat and its prey species. The Indian government has planned a five-year "Project Snow Leopard" modelled on the successful "Project Tiger" beginning in 1990 and aimed at securing the recovery of the species. The species breeds well in captivity. A Species Survival Plan run by the AAZPA exists to manage the breeding programme for about 200 of the approximately 450 Snow Leopard currently held around the world (Wharton, 1991).

Sources:

Fox, J. L., Sinha, S. P., Chundawat, R. S. & Das, P. K. (1991). Status of the Snow Leopard Panthera uncia in Northwest India. Biol. Conserv. 55, pp 283-298.

Jackson, P. (1987). Symposium on the Snow Leopard. Cat News 6: 14-17.

Schaller, G. B. (1988). The Snow Leopard in China. Cat News 9: 29 pp.

IUCN (1978). Snow Leopard Red Data Sheet. IUCN, Gland, Switzerland, and Cambridge, U.K.

Wharton, D. (1991). Glasnost among zoos. p N5 in: Wildlife Conservation. March / April 1991.

Family Mustelidae

ORIENTAL SMALL-CLAWED OTTER
Status: INSUFFICIENTLY KNOWN
Aonyx cinerea (Illiger, 1815)CITES: Appendix II

HB 41-64cm; T 25-35cm (Macdonald)

The Asian Small-clawed Otter is the smallest of the world's otters, rarely weighing more than 5 kg. The coat is coloured shades of brown, the throat being lighter (white or grey). Asian Small-clawed Otters have unusual hand-like front paws with increased tactile sensitivity and reduced webbing, which they use to forage for their prey of crustaceans, molluscs and small fish. A gregarious species, it is often seen in large groups of up to 15 animals, and captive studies suggest that these groups are composed of a dominant breeding pair and their offspring from successive litters. The species is found from Palawan (Philippines) through Indonesia, southeast Asia, southern China, and westwards through the Himalayan foothills of Bangladesh, Bhutan and Nepal. A disjunct population occurs in southern India. This species is widespread and common in parts of its range, but its distribution is poorly known or unknown in several countries. Threats include illegal hunting for domestic markets, fish traps also catch and drown otters. Special management practices are required to ensure the survival of the species in particular habitats, for example paddyfields, where it plays a valuable role in the control of pests. There is need to develop new protected areas for the species, especially in the mangrove areas of Peninsular Malaysia. It has been kept in captivity. A reintroduction programme for the species should be considered in Hong Kong.

Sources:

Foster-Turley, P., Macdonald, S. and Mason, C. (Eds.) (1990). Otters, An Action Plan for their Conservation. IUCN/SSC Otter Specialist Group. IUCN: Gland, Switzerland. 126 pp.

Macdonald, D.W. (1984). The Encyclopaedia of Mammals. Unwin Hyman, London, UK. 895 pp.

SMOOTH-COATED OTTER
Status: INSUFFICIENTLY KNOWN
Lutra perspicillata I. Geoffroy, 1826CITES: Appendix II

HB 65-75cm; T 40-45cm; W 7-11kg (Burton and Pearson, 1987)

Smooth otters are so-named for their shorter, smoother coats, as compared to the similar sized sympatric Lutra species. From captive studies this otter appears to be monogamous and is usually seen year-round in small family groups. Although little is known of the ecology of the species, it is known to occur in a variety of Asian habitats, from mangroves to freshwater wetlands and large forested rivers. The smooth-coated otter is distributed throughout southern Asia from Indonesia through southeast Asia, and westwards through southern China and India, with an isolated population in Iraq (L. p. maxwelli). In most of its range it is sympatric with the Asian Small-clawed Otter and sometimes the other Lutra species. Of all the Asian otters, this one seems to be the most common through most of its range.

Sources:

Burton, J.A. and Pearson, B. (1987). Collins Guide to the Rare Mammals of the World. Collins, London, UK. 240 pp.

Foster-Turley, P., Macdonald, S. and Mason, C. (Eds.) (1990). Otters, An Action Plan for their Conservation. IUCN/SSC Otter Specialist Group. IUCN: Gland, Switzerland. 126 pp.

NILGIRI MARTEN
Status: VULNERABLE
Martes gwatkinsi (Horsfield, 1851)CITES: Appendix III

The Nilgiri Marten is dark brown with reddish brown foreparts, it is smaller than M. flavigula, but its coat is similar. Found in southern India; specifically the southern parts of Western Ghats, and associated hill ranges from approximately 13°N, south to the hills of Travancore. Reports indicate that it survives in forest patches on mountain summits, mainly semi-evergreen and evergreen forests, it is sometimes encountered in coffee and cardamon plantations. The marten is legally protected, but it is occasionally persecuted as a pest by bee-keepers. The species is known to occur in the Eravikalum-Rajamalai National Park (9,700 ha) in Kerala and in Brahmagiri Wildlife Sanctuary (18,100) in Karnataka, and possibly in several others. Field surveys are required to locate remaining populations of the subspecies and to determine whether the existing reserves give adequate protection to M. gwatkinsi and its habitat.

Source:

Schreiber, A., Wirth, R., Riffel, M. and Van Rompaey, H. (1989). Weasels, Civets, Mongooses and their Relatives, An Action Plan for the Conservation of Mustelids and Viverrids. IUCN/SSC Mustelid and Viverrid Specialist Group. IUCN: Gland, Switzerland. 100 pp.

Family Ursidae

LESSER OR RED PANDA
Status: VULNERABLE
Ailurus fulgens F. Cuvier, 1825CITES: Appendix II

HB 51-63.5cm; T 28-48.5cm; W 3-4.5kg (Nowak 1991)

The Lesser or Red Panda resembles a racoon in size and form, with reddish-brown fur and a ringed tail, and weighs on average around 5 kg. This mainly nocturnal species inhabits Himalayan temperate forests with bamboo thicket understories at relatively high altitudes (between 2,200 and 4,800 m), distributed from Nepal to China (Yunnan and Sichuan provinces), including Burma, Bhutan and India. The large scale habitat destruction is threatening the species, especially in the west of its range; forests are cut for wood and cleared for agriculture. It has been traded for its fur and the pet trade in the past. There are taxonomic difficulties in classifying the species, whether it is a raccoon, bear, or to be grouped separately with the Giant Panda. The Red Panda in China is sympatric with the Giant Panda, both feeding on bamboo (although a variety of other foods are eaten including some animals and fruits); if the habitat of the popular Giant Panda is conserved, this will also aid the conservation of the Red Panda (the Chinese government has created 12 reserves for the Giant Panda). It is protected in India and Nepal. In response to the scarcity of the species, an international breeding programme was initially established in more than 30 zoos; it is now a popular exhibit in zoos.

Sources:

Anon. (1991). Cut cheese production to save pandas. Oryx Vol. 25 (4), p 190. (refers to Conservation Biol. (1991) Vol. 5, pp. 196 - 202).

Johnson, K. G. (1990). Mystery of the Other Panda. In: International Wildlife. Vol.20, No.6. Nov/Dec 1990: 30-33.

Nowak, R.M. (1991). Walker's Mammals of the World, 5th edn. The John Hopkins University Press, Baltimore and London. 1629 pp.

Roberts, M. S. (1984). Ailurus fulgens. Mammalian Species. No. 222, Nov 1984: 1-8.

SUN BEAR
Status: VULNERABLE
Helarctos malayanus (Raffles, 1821)CITES: Appendix I

HB 1-1.4m; T 3-7cm; WT 27-65kg (Burton and Pearson, 1987)

The Sun Bear was formerly widespread in the lowland forests of South-East Asia from India eastward through Myanmar, southern China, Laos, Viet Nam, Thailand, Malaysia, and Indonesia. The species may now be extinct in India, Bangladesh and China, and the range is shrinking in most areas. Sun Bears inhabit protected areas in Thailand, Malaysia, Indonesia and possibly the Siju Sanctuary in India. In Thailand the species is called the "dog bear" because of its small size and short hair; it is the smallest of the eight bear species. It is mainly nocturnal, inhabits forest, preferring high branches and diet includes termites and honey. The Sun Bear is threatened by trade both for pets and to provide various products; bears are in demand for their meat and gallbladders to supply the expanding illegal trade market. Habitat destruction is another major threat, often caused by conversion of lowland areas to plantations. The Sun Bear is protected by law in China (a "first class conserved animal"), Thailand, and Indonesia. Legislation to conserve the species is poorly enforced, if at all, in most countries of its range. The Sun Bear is the least well-known of all bears, with a very bleak future due to the lack of management and understanding, coupled with habitat destruction and poaching (Servheen, 1990 and Mills, 1991). The Sun Bear is currently being studied in Sabah, Malaysia, involving officials in Sabah (Servheen, 1991).

Sources:

Burton, J.A. and Pearson, B. (1987). Collins Guide to the Rare Mammals of the World. Collins, London, UK. 240 pp.

Mills, J. (1991). "I want to eat Sun Bear". International Wildlife. Vol.21, No.1. Jan/Feb 1991: 39-43.

Servheen, C. (1990). The status and conservation of bears of the world. International Conference on Bear Research and Management Monograph Series No. 2. 32pp.

Servheen, C. (1991). Update on Ongoing Projects Worldwide. IUCN/SSC Bear Specialist Group Newsletter. No.2, Feb. 1991. 9 pp.

SLOTH BEAR
Status: VULNERABLE
Melursus ursinus (Shaw, 1791)CITES: Appendix I

HB 140-170cm; SH 65-85cm; WT 125-145kg

The Sloth Bear has long dark fur highlighted with white on the muzzle, the tips of the paws and in a V-shaped breast patch. It is nocturnal except in the remoter areas when it may be active on cooler days. At night it searches for fruits and insects and will also take carrion. It is found throughout forested areas south of the Himalayas, in India, Sri Lanka, Bangladesh, Nepal and Bhutan. It predominantly inhabits low elevations, its optimum habitat being the dry deciduous forests of central India. The species was common in most forested areas of India up until the 1950s. In 1975 the population of Sloth Bears in India and Sri Lanka was estimated at fewer than 10,000 and declining (Jaffesson, 1975). Major threats include forest cutting and intrusion of settlements into bear habitat, which can lead to crop raiding incidents. It has a low reproductive and recruitment rate in the wild, and is therefore extremely vulnerable to hunting. Bear species in India are killed for the sale of parts to Bangladesh, and the Sloth Bear is no exception; illegal hunting is also a problem in Sri Lanka. The gall bladder is traded for its reputed medicinal value. In 1990 the 62 kg of gall bladders imported into Japan from China was estimated to have a retail value in Japan of 3.9 million U.S. dollars (Anon, 1990). The Sloth Bear is classified as endangered by the Indian government; the species is protected and sale of parts and exportation is illegal. Sloth Bears have been reported from 13 protected areas in India; 3 sanctuaries have been specifically created for the species. It is protected in Sri Lanka, where it occurs in the low country dry zone, including protected areas such as Rhuna National Park. In Nepal it is partially protected by law and occurs in protected areas such as the Chitawan National Park (Santiapillai, 1989). There is little information available on the species and further research is required (Servheen, 1990). A two-year radio-telemetry research study in the Chitawan National Park began in 1990 to examine bear movements, habitat use and food habits (Servheen, 1991). The long-term survival of the species can only be certain in the few larger, more remote protected areas. Surveys are required to determine areas where additional protection is needed urgently.

Subspecies:
Melursus ursinus inornatus Pucheran, 1855Sri Lanka
Melursus ursinus ursinus Shaw, 1791India

Sources:

Anon (1990). IUCN/SSC Bear Specialist Group Newsletter. No.1, January 1990: 2.

Jaffesson, R.C. (1975). Melursus ursinus: Survival Status and Conditions. Ms. No.3. Washington D.C., USA. 15 pp.

Prater, S.H. (1971). The Book of Indian Mammals. Bombay Natural History Society. pp. 139-140.

Santiapillai, A. and Santiapillai, C. (1989). Status, Distribution and Conservation of the Sloth Bear (Melursus ursinus) in Sri Lanka. WWF-Indonesia Programme: Bogor, Indonesia. 6 pp.

Servheen, C. (1990). The status and conservation of bears of the world. International Conference on Bear Research and Management Monograph Series No. 2. 32pp.

Servheen, C. (1991). Update on Ongoing Projects Worldwide. IUCN/SSC Bear Specialist Group Newsletter. No.2, Feb. 1991. 9 pp.

ASIATIC BLACK BEAR
Status: VULNERABLE
Selenarctos thibetanus (G. Cuvier, 1823)CITES: Appendix I (= Ursus thibetanus)

HB 1.2-1.8m; T 6.5-10cm; WT 90-180kg (Prater, 1971; Burton and Pearson, 1987)

The Asiatic Black Bear has a coat of smooth black fur except for the distinct 'V' shape of white fur on the chest. Its black claws easily distinguish it from the Sloth Bear (Melursus ursinus), which has white claws. It feeds upon a wide variety of vegetation, fruits, insects and sometimes small mammals. It inhabits forested uplands, mountains and tropical moist forest below alpine elevations in much of southern Asia. The former range extended from Afghanistan and Pakistan in the west, across northern India, Indochina and southern China, and to northeastern China, eastern Russia, the Korean Peninsula and Japan. The species may now be extinct in Afghanistan, and is very close to extinction in South Korea and Bangladesh. It is threatened by poaching, opportunistic killing and agricultural expansion throughout its range. The species is in demand in Thailand as pets and for products; for example, in 1989 single servings of bear paw soup sold for several hundred U.S. dollars. The export of bear gall bladders has been a major problem in China, the trade provides a large source of foreign capital as gall bladders can fetch high prices. Japan continues to trade in Asiatic Black Bear products through an alteration of their CITES agreement. Specific legislation and efficient implementation is needed to protect bears from hunting. The species could become extinct throughout most of its range in the near future unless there are stricter controls on the trade in bear products. Asiatic Black Bears are protected by legislation in South Korea (although bears are imported for traditional medicinal products), and in eastern Russia. The species is marginally protected by law in Thailand and China. They are known to inhabit Indian protected areas, and hunting has been regulated in India since 1972 (Servheen, 1990). There are plans to study the natural history of the species in reserves in Taiwan, and a primary census is planned for Sichuan, China in 1991 (Servheen, 1991).

Sources:

Burton, J.A. and Pearson, B. (1987). Collins Guide to the Rare Mammals of the World. Collins, London, UK. 240 pp.

Prater, S.H. (1971). The Book of Indian Mammals. Bombay Natural History Society.

Servheen, C. (1990). The status and conservation of bears of the world. International Conference on Bear Research and Management. Monograph Series No. 2. 32pp.

Servheen, C. (1991). Update on Ongoing Projects Worldwide. IUCN/SSC Bear Specialist Group Newsletter. No.2, Feb. 1991. 9 pp.

Family Viverridae

JERDON'S PALM CIVET
Status: VULNERABLE
Paradoxurus jerdoni Blanford, 1885CITES: Not Listed

HB 59cm; T 52cm; WT 3.6kg (Macdonald)

Jerdon's Palm Civet is endemic to the evergreen rain forest belt of southwest India, distributed in Western Ghats and associated hill ranges, possibly as far north as Uttara Kannada district, Karnataka. The species is known to inhabit humid forest and coffee plantations. The coat is dark brown to black with grey speckling. The species has been kept in captivity (1969). The occurrence in several protected areas is confirmed, and considered likely in several others. Protection of the remaining forests throughout the Western Ghats is required to conserve the species. A field survey in the Western Ghats to identify areas where populations of the species still survive and what their conservation needs are is required.

Sources:

Macdonald, D.W. (1984). The Encyclopaedia of Mammals. Unwin Hyman, London, UK. 895 pp.

Schreiber, A., Wirth, R., Riffel, M. and Van Rompaey, H. (1989). Weasels, Civets, Mongooses and their Relatives, An Action Plan for the Conservation of Mustelids and Viverrids. IUCN/SSC Mustelid and Viverrid Specialist Group. IUCN: Gland, Switzerland. 100 pp.

Order CETACEA

Family Delphinidae

IRRAWADDY DOLPHIN
Status: INSUFFICIENTLY KNOWN
Orcaella brevirostris (Gray, 1866)CITES: Appendix II

The Irrawaddy Dolphin is a warm-water species found only in the tropics and subtropics, mostly in shallow coastal waters. Nevertheless some animals probably live all their life in freshwater and they occur in many major river systems, reputedly as far as 1,300 km from the sea. This species is restricted to the coasts and rivers of Southeast Asia and northern Australia, from the Bay of Bengal to north Australia. Population figures are scarce but there is no indication that this species is currently under threat, although its habitat is vulnerable to dams and other human disturbance. Because of their coastal and riverine habitat, Irrawaddy dolphins often come in contact with man. They are killed for food in only a small part of their range, but often become accidentally entangled in fishing nets and even in anti-shark nets in northern Australia. Fishermen in Cambodia and Vietnam are said to regard them as sacred and use them to drive fish into nets. These dolphins have been kept in several captive facilities in Southeast Asia, where breeding has been successful.

Source:

Martin, A.R. (1990). Whales and Dolphins. Salamander Books Ltd.: London.

Family Platanistidae

GANGES RIVER DOLPHIN
Status: VULNERABLE
Platanista gangeticaCITES: Appendix I

The Ganges River Dolphin inhabits the Ganges, Brahmaputra, Karnaphuli and Meghana Rivers and their tributaries in India, Bangladesh, Nepal and Bhutan. The species prefers deep water and migrates seasonally upstream as the water level rises. It feeds mainly on fish, shrimps and molluscs, relying heavily on echolocation and touch to find its prey due to its poor eyesight. Formerly abundant, but now reduced to approximately 5,000 individuals (3-4,000 in India), the species is threatened by pollution, dams, mining and directed and incidental catch. The dolphins of the Ganges system are now divided into small isolated subpopulations. The Ganges River Dolphin is protected by nature conservation laws in Nepal and India. Creation of protected areas could prove effective in conserving the dolphin and suggestions have been made for specific sites. Surveys of populations would also assist conservation efforts, as would campaigns promoting awareness of the species amongst the local people.

Source:

Perrin, W., Brownell, J., Kaiya, Z. and Jiankang, L. (1989). Biology and Conservation of the River Dolphins. Proceedings of the Workshop on Biology and Conservation of the Platanistoid Dolphins, at Wuhan, P.R. of China. IUCN: Gland, Switzerland.

Order PERISSODACTYLA

Family Equidae

ASIATIC WILD ASS
Status: VULNERABLE
Equus hemionus (Pallas, 1775)CITES: Appendices I & II

Also known generically as the Kulan or Onager, the Asiatic Wild Ass was once abundant over a large part of arid and semi-arid E. Europe and Asia from Bulgaria and Roumania in the west to Manchuria in the east. Five subspecies are recognised. The Syrian Wild Ass (E. h. hemippus) is believed to be Extinct and the Indian Wild Ass or Khar (E. h. khur) is listed as Endangered. These two subspecies are dealt with separately. The whole species is collectively classed as Vulnerable and listed on CITES Appendix II. The Dzigettai or Mongolian Wild Ass (E. h. hemionus) is now mainly restricted to high semi-desert grassland, river margins and gorges in southern Mongolia, where there is a population in the Great Gobi National Park. It ranges eastwards to 106oE and is also be found in Sinkiang, China, where the population was estimated at approximately 1,000 in 1988 (Clark 1991). Dzigettai weigh approximately 290 kg, are grazers, and feed mainly on cereal species and wormwood. Females and young live in breeding herds with a single adult male: other males form bachelor groups. Herds may aggregate on good pastures or during winter. The ecology of the other subspecies of Asiatic Wild Ass is believed to be similar. The Dzigettai is fully protected by law in China and Mongolia and is listed on Appendix I of CITES. Scattered populations of the Kiang (E. h. kiang) occur in similar semi-desert habitats on the high Tibetan plateau and in Protected Areas in adjacent parts of both India and Pakistan. The Kiang is fully protected under Chinese law. The Onager (E. h. onager) persists only in and around Protected Areas in northern Iran and Turkmenistan in the USSR, possibly migrating into Afghanistan. It is fully protected by law in both Iran and the USSR, and is the only subspecies to be breeding well in captivity. The Asiatic Wild Ass has been in decline for over 2000 years, mainly due to exclusion from forage and water supplies by human settlement and domestic stock grazing, although hunting and disease have also contributed. Provided Protected Areas continue to be policed effectively, the Asiatic Wild Ass may easily be saved, since it now occupies habitats that are sub-optimal for humans and their livestock.

Sources:

IUCN (1979). Asiatic Wild Ass Red Data Sheet. IUCN, Gland, Switzerland and Cambridge, U.K.

Clark, B. (1991). Wild Assumptions. BBC Wildlife, Vol. 9 No. 10, p 737, August 1991.

INDIAN WILD ASS
Status: ENDANGERED
Equus hemionus khur (Lesson, 1827)CITES: Appendix I

The Indian Wild Ass, Khar or Gorkhar, inhabits remote parts of the Little Rann of Kutch, a 259,000 ha. salt plain along the Indian / Pakistan border. The Asses that live in the Qatruaye Basin, a restricted area about 160 km east of Shiraz in Iran may also belong to this subspecies. The Khar inhabits hills during the monsoon, and in the dry season descends and forages along the edge of the Little Rann, where it frequently raids cultivated fields during darkness. The main threats to the Khar are overgrazing by livestock; persecution by farmers in retaliation for crop-raiding; and contraction of diseases from domestic horses. The Khar was given full legal protection in 1952, and the entire Little Rann of Kutch was declared a Wildlife Sanctuary in 1973. The last known census indicated that the population had increased from 700 in 1976 to 2,000 in 1983 (D'Souza 1983). Conservation measures necessary include the prohibition of firearms, salt extraction and grazing of domestic stock in the Sanctuary, and a more strictly enforced campaign of vaccination against horse sickness in the surrounding settlements.

Sources:

D'Souza, E. (1983). In litt. to WCMC

IUCN (1979). Asiatic Wild Ass Red Data Sheet. IUCN, Gland, Switzerland and Cambridge, U.K.

Family Rhinocerotidae

SUMATRAN RHINOCEROS
Status: ENDANGERED
Dicerorhinus sumatrensis (Fischer, 1814) CITES: Appendix I

At 800 kg the smallest member of the family, the Sumatran or Asian Two-horned Rhinoceros inhabits montane rain forest in South East Asia. A solitary species, it is primarily a browser, feeding on fruit, leaves, twigs and bark. Its sparse covering of long hair - in sharp contrast to the other hairless species of rhinoceros - has earned it the soubriquet of Woolly Rhinoceros. The original distribution stretched from Bhutan and eastern India through Burma, southern China, Thailand, and Malaysia to Sumatra and the island of Borneo (with unconfirmed reports of its presence in northern Laos, Cambodia and western central Vietnam). Only 500 - 900 Sumatran Rhinoceros are believed to remain, mostly in scattered inviable populations in Burma, Thailand, the Malay Peninsula, Sumatra and Borneo (Khan 1989). The decline has been caused mainly by hunting to provide horns and other parts of the carcase for traditional Oriental medicines. In 1990 Asian rhino horn was reaching a wholesale value of up to US$54,000 per kg in Taiwan, partly because wealthy individuals were buying it not only to cure ailments but as an investment, since as wild stocks decline the price is expected to leap still higher (Bradley Martin and Bradley Martin 1991). Habitat destruction due to logging and agricultural development is however increasingly affecting the species - particularly in Sumatra which harbours the bulk of the remaining wild population and is under intensive development by Indonesia's transmigration programme. The Sumatran Rhinoceros is extremely sensitive to human presence, and readily abandons areas in which it has been disturbed. Conservation measures proposed include the capture of "doomed" animals from the wild to further develop the small captive population (24 were held in collections in the U.K., the U.S.A., Malaysia, Thailand and Indonesia in mid-1991 (Foose, 1991)); the concentration of in situ conservation efforts on the 7 - 9 wild subpopulations considered to be reasonably viable in the long-term out of the known total of 35; gazetting of new Protected Areas (e.g. Kerinci Seblat in Sumatra may now (5/5/92) have been gazetted - check) to supplement those already containing the species; improved enforcement of existing legislation against poaching and trade in rhinoceros products; and the financing of surveys in Kalimintan (Indonesia), Thailand and northern Burma to determine whether appreciable populations of Sumatran Rhinoceros survive there.

Sources:

Bradley Martin, C. and Bradley Martin, E. (1991). Profligate spending exploits wildlife in Taiwan. Oryx Vol 25 No. 1: 18 - 20.

Foose, T. (1991). Summary of Sumatran Rhino Captive Programs - 1984 to 1991. Around the Horn, Vol. 2, No. 1: 9.

Khan, M. K. M. (1989). Asian Rhinos An Action Plan for their Conservation. IUCN/SSC Asian Rhino Specialist Group. IUCN/WWF Gland.

INDIAN RHINOCEROS
Status: ENDANGERED
Rhinoceros unicornis Linnaeus, 1758 CITES: Appendix I

Also known as the Great or Greater One-horned Rhinoceros, the Indian Rhinoceros once ranged across the entire northern part of the Indian sub-continent from Pakistan through parts of Nepal, Bhutan and Bangladesh to the Indo-Burmese border. It may also have existed in Burma, southern China and Indochina. A large (2,000 - 4,000 kg) solitary species which mainly inhabits floodplain grasslands alongside rivers and lakes but is also found in drier forest areas in Nepal, the Indian Rhinoceros is primarily a grazer. The species has been eliminated in most areas by hunting for its horn, and habitat loss to agriculture and stock grazing. It is now confined to isolated pockets of its original range in north-eastern India, Nepal, and Pakistan, and the bulk of the surviving animals occur in Protected Areas. Under strict protection by the Indian and Nepalese wildlife authorities numbers have increased from an estimated 1,000 - 1,100 in 1977 (WCMC 1978) to 1,700 in 1987 (Khan 1989) and an estimated 1,950 in 1991 (Bradley Martin and Vigne 1991). The majority of these - 1,510 - inhabit the Indian state of Assam, where approximately 1,250 are found in the Kaziranga National Park (Vigne and Bradley Martin, 1991). Animals have successfully been translocated to re-establish populations in areas where they had been exterminated. Although the Indian Rhinoceros is the least threatened of the Asian species, poaching pressure has recently intensified: 58 were killed in Assam in 1990, including at least 7 which were electrocuted by poachers using high voltage wires that run through the Kaziranga and Pobitara sanctuaries (Anon. 1991). Encroachment by agriculturists on Protected Areas continues; and invasion by the exotic plants Mikania scandens and Eichhornia crassipes is causing problems in Assam and West Bengal by smothering and preventing growth of native grasses, producing serious loss of rhino food resources. Conservation measures proposed include the maintenance of a wild population of at least 2,000 rhinos; concentration of field efforts on the six sanctuaries which contain reasonably viable wild populations (i.e. > 100 animals), namely Kaziranga, Manas, Orang and Dudhwa in India, and Chitawan and Bardia in Nepal; clampdown on illegal trade in rhino products; and an increase in the size of the current captive world herd from 75 to 150 (Khan, 1989).

Sources:

Anon. (1991). Rhinos electrocuted. Oryx 25: 65.

Bradley Martin, E. and Vigne, L. (1991). The horn quintet. BBC Wildlife, Vol. 9. No. 5. pp 356-357.

Khan, M. K. M. (1989). Asian Rhinos An Action Plan for their Conservation. IUCN/SSC Asian Rhino Specialist Group. IUCN/WWF Gland.

Vigne, L. and Bradley Martin, E. (1991). Assam's rhinos face new poaching threats. Oryx, Vol. 25 (4): 215 - 221.

Order PROBOSCIDEA

Family Elephantidae

INDIAN ELEPHANT
Status: ENDANGERED
Elephas maximus Linnaeus, 1758CITES: Appendix I

The Indian, Asiatic or Asian Elephant once occurred in a variety of lowland and montane forest and savanna habitats up to 3,600 m from the Tigris and Euphrates basin in Syria and Iraq, through Asia south of the Himalayas and north into China. Today it occupies scattered fragments of its former range in 13 countries: Bangladesh, Bhutan, Burma, Cambodia, China, India, Indonesia, Laos, Malaysia, Nepal, Sri Lanka, Thailand and Vietnam. Only between 34,000 and 54,000 - one tenth of the number of African elephant - survive in a land area of just 500,000 km2 (Santiapillai & Jackson 1990). The population is concentrated in India (17 - 22,000 head) and Burma (3,000 - 10,000 head). Indian Elephants weigh 4 - 5 tonnes and eat up to 150 kg of vegetation each day. Grass forms the principal component of the diet, and its availability determines seasonal elephant movements. The species is gregarious: females and young live in matriarchal family units of approximately 6 animals, while adult males live singly or in small groups and have no permanent ties with the females. The Indian Elephant adapts well to domesticity, and has been used for centuries as a draught animal in forest industries, warfare, and ceremonials. It breeds well in captivity, and there are currently approximately 16,000 tame elephants, mostly employed in the timber industry in Burma, India, Indo-China and Thailand. Most of these have been captured from the wild. The Indian Elephant is legally protected in most countries of its range and occurs in numerous Protected Areas, but poaching for ivory (and in Burma for hides which are exported to Thailand) continues. Although since Roman times exploitation for ivory was the main factor in the Indian Elephant's gradual decline, the species is not currently greatly threatened by the international ivory trade. Unlike the African Elephant, only males have tusks, and many males are tuskless, so even intensive poaching for ivory leaves a nucleus of breeding females. Instead the major problem faced by the Indian Elephant in the 20th century is habitat destruction and fragmentation of populations. It is now almost totally excluded from lowland habitats by human occupation, and in hilly areas where it is compressed into small pockets of remnant forest, conflict with surrounding agriculturists occurs. Crop-raiding is frequent and there are often human deaths. In India alone about 200 people are killed every year. Proposed conservation measures include the stricter enforcement of national laws and CITES regulations; creation of new Protected Areas and the establishment of large Managed Elephant Reserves where human activities compatible with conservation would continue, but elephants would have priority in management decisions; maintenance of forest corridors to facilitate migration between Protected Areas; establishment of compensation schemes for crop losses and measures to counteract and prevent crop-raiding, including the planting of barriers of crops unattractive to elephants such as tea and oilseeds, and the construction of non-lethal electric fencing; translocation of elephants, both away from nuisance areas and to maintain genetic variability of small populations; and ecological research and public education programmes. In India Project Elephant (along the lines of Project Tiger) was scheduled to be launched in 1990-1991.

Source:

Santiapillai, C. & Jackson, P. (1990). The Asian Elephant: An Action Plan for its Conservation. IUCN/SSC Asian Elephant Specialist Group. IUCN, Gland, Switzerland.