The definition of a shoal is a
'group of fish which remain together for social reasons,
including feeding, reproduction or resting and can
comprise of only a few fish to millions in the open
sea'.
In every day conversation we often class shoaling
and schooling as one behavioural trait. This is not
the case, although both mean that the fish stay in one
group, they do have different characteristics in the
way they form the shoal or school.
Shoals contain fish which swim together with no
formation pattern or in any synchronised way. They
have a varying distance between them and their
neighbouring fish.
Schools, on the other hand contain fish which swim in
an organised and planned way. They tend to swim at
the same speed and keep close together. Shoals of
fish that are on the move almost always form schools.
ADVANTAGES
The main advantage of shoaling is that of
protection from predators. The phrase 'two heads are
better than one' is well known and applies in this
case. Except it is more likely to be 'hundreds of
heads are better than one'. Shoaling multiplies the
number of eyes and the directions in which they are
looking for predators. This increases the probability
of a predator being seen before it attacks. Fish alert
each other by moving in a certain way. For example,
the European minnow performs a movement called the
'skitter' A skittering fish alerts the other members
of the shoal that danger has been spotted. Co-ordinated
movements of shoals are thought to deter and even in
some cases prevent predator fish from attacking if the
shoal is large enough.
Shoaling also increases the group's chance of
successful migration. Many species migrate and
experiments have proved that individuals are far less
successful in negotiations than large shoals. Another
advantage to fish when they shoal is that it is
thought that as a group they may be able to reduce
drag and therefore be able to swim more efficiently.
I say that this is a thing which is 'considered' an
advantage because there has actually been no
experimental proof. However, it works in theory.
DISADVANTAGES
Living in a large group means that there is heavy
competition for food. Living in a shoal means that
resources have to be stretched to suit everyone and
when such a situation occurs there in inevitable
tension.
SHOALING IN A TANK
You would expect fish to stop shoaling when in a
tank because it is largely a predator free zone. It is
common knowledge, though, that fish are stupid and
they don't know if there isn't a predator in the tank.
They are always expecting to find a predator waiting
behind the next leaf! So, if a fish is classed as a
shoaling fish it is best to keep them in a shoal even
in a tank, because without other members of its
species it will feel vulnerable and will become
stressed due to this. Fish have been known to group with
other species, for example a Guppy may shoal with a
minnow. But given the choice, a fish will always
shoal with its own kind.
When attacked, fish have many co-ordinated
manoeuvres that they may perform. Some fish simply
perform what is known as a 'U-turn' - this means they
divide into two and pass the predator on either side.
This is performed in the hope the predator will be
confused by the fact his target has split and passed
him on either side. Another escape response it to
divide in half, again, and to swim off in opposite
directions. This movement is also hoped to confuse
the predator by the fact that he will watch his
intended target move in opposite directions and become
confused. Perhaps the most attractive movement is for
the shoal to 'explode'. When fish 'explode' they all
try to escape in opposite directions. However, if the
shoal is not quick enough to regroup they face the
danger of being picked off slowly. The last of the
manoeuvres is for fish to form into a tight ball.
However, may people consider this to be a 'selfish'
act by the shoal as only the fish in the middle are
being protected by those on the outsides.
Shoaling is thought to lessen the chance of being
eaten. It is true that a predator will take longer to
find one big group than many scattered individuals.
However, once a predator has found a fish it has the
same probability of being eaten whether it is in a
large shoal or on its own.
(Adapted from Practical
Fishkeeping, written by Dr Gordon Watson, October 1999
issue)