SLOVENIAN ROOTS:  WHAT GENES REVEAL

(Korenine Slovencev:  Kaj razodevajo geni)

(J. Skulj (P. Eng.) The Hindu Institute of Learning, Toronto, Canada – 2004-3-30)

(Translation from Slovenian by Lillian Centa -- 8/04)

Summary:
In the year 2003, Malyarchuk et al. in collaboration with a Slovenian, Dr. K. Drobnic (M), published their research paper “Mitochondrial DNA Variability in Bosnians and Slovenians.”  The results made it possible for Slovenians to compare themselves, genetically, with other nations through genetic markers (haplogroups) inherited from the mother – mtDNA. – as was possible from the year 2000 with genetic markers from the Y-chromosome inherited from the father (SS).  From literature, I summarized the genetic data of various nationalities that would be of interest to Slovenians.  It is evident from the comparison that 90% of Slovenians have the mtDNA genetic markers that originated before or during the last ice age; whereas, only 75% of the 4,000 to 5,000 year old skeletons, excavated in the Basque country have genetic markers that are now that old; and the Basques of today have 70% of the genetic markers that originated before or during the last ice age.  Only 1% of the Slovenians have genetic groups 6,000 to 13,000 years old; their other genetic groups are all much older.

Archaeological finds:
Archaeologists maintain that man has lived in Europe for at least 800,000 years.  This early man was given the name Neanderthal, after the Neanderthal gorge near Dusseldorf in Germany where his bones were first discovered.  Even older bones, resembling human, believed to be 1.75 million years old, were discovered by archaeologists in Dmanisi in today’s Republic of Georgia(G).  The oldest remnants of tools were found by archaeologists in 1995 in Spain.  They estimate that these tools are about 789,000 years old.

Slovenia is also an interesting place for archaeologists.  The oldest trace of pre-historic man in Slovenia was found in Divje babe (“wild women”) above the Idrijca River in 1996, when a whistle made of bone was found dating from the period prior to 45,000 years ago, which they ascribed to the Neanderthal man.(TM)  As an engineer, I was surprised by the elaborate whistle on account of the workmanship involved in its production.  There is evidence that a drilling technique was used in its manufacture.  Of the four holes, two are not damaged, but all four show signs of drilling, since they are very symmetrical.  It is possible that even then, the Neanderthals used the same drilling techniques that were used by the Eskimos who used a bow and stone-tipped arrows for drilling, even after the arrival of Europeans.  Another sign of the developed primitive technological skill is the oldest, more than 5,000 years old, wooden wheel with an axle that was found in the swamplands near Ljubljana.  To make this wheel and transverse axle, the craftsmen had to use axes and probably also saws.  It is interesting that the wheel did not turn on the axle, but the axle with the wheel turned on bearings, similar to a system used, even today, on modern railway cars.  We are able to take the wheel with the integral axle as evidence of a relatively highly developed woodworking technology and also as a sign of metallurgy, since a mold for copper axes was found not far from the site of the pre-historic wheel.

Recently, in Croatian Zagorje, at Vindija and Velika Pecina, bone and stone tools were found.  It is believed that they belonged to Neanderthals who may have lived there more than 29,000 years ago.  Nearby, tools of stone and bone were also found, characteristic of modern man.  It may be that the Neanderthals made the tools themselves or acquired them through trade.  It was previously believed that the Neanderthal man became extinct more than 34,000 years ago.  This new evidence compelled scientists to change their theories.  Fred Smith, an anthropologist from Northern Illinois University, says that it is possible that the Neanderthal man and modern man lived in close proximity in central Europe for many thousands of years.  Erik Trinkhaus, an anthropologist from Washington State University, thinks that modern man in some places superseded the Neanderthals, elsewhere, he intermarried with them.  Between these two races, Trinkhaus sees very small distinctions.  In 1999, he announced that he found, in Portugal, the bones of a child from 24,500 years ago, which showed characteristics similar to those of both the Neanderthal man and of modern man.  Trinkhaus and Smith thus raised the new, controversial, hypothesis that Neanderthal man and homo sapiens intermarried.(NP)

Genetic data:
Geneticists such as Ridley and others, who are extremely interested in the study of human genes, have discovered that for the last four billion years, the human genome has been recording significant events in our biography.  There are genes with the help of which it is possible to track man’s migration during the last thousands of years.(RM)  Gutierrez and others theorize about the possibility, that some of the Neanderthal genes are also represented in the genetic structure of Europeans.  Gutierrez proved that genetically some Africans are more distant from Europeans, than three Neanderthals to whom they were compared using genetic technology.(G)  Calafell has observed that in today’s populations, there are more genetic distinctions in populations whose ancestors had a possible contact with Neanderthals in prehistoric times.(C)  Thus some geneticists also agree with anthropologists.

The year 2003 was a new milestone for Slovenian historical research, when a Russian geneticist, Dr. Boris A. Malyarchuk published his paper, “Mitochondrial DNA Variability in Bosnians and Slovenians”.  This paper shows the genetic profile of Slovenians based on genetic markers inherited on the mother’s side-the mtDNA.  This is a big step forward, since we now have the possibility of genetically comparing Slovenians with other nations, not only on the basis of genetic groups from the Y-chromosome inherited from the father, but also on the basis of ancestral groups inherited on the mother’s side.  This will have far-reaching consequences for the re-writing of Slovenian prehistory, because this is additional, unbiased evidence, that Slovenians are an ancient people, living in their own land continuously for thousands of years.

In the tabulated list below, I took the mtDNA haplogroup data from literature of other nations and their ages, and compared them with Slovenians.  Included in this comparison, in addition to contemporary people, are also genetic groups from people, who 4,000 to 5,000 years ago, lived in the Basque region of Spain.(BSK1)  If we statistically analyze data from the tabulated list and consider the weighted averages, we see that the average genetic age of Europeans is 26,710 years.  With Slovenians, the average calculated age of mtDNA genetic groups is about 750 years older at – 27,460 years.

MtDNA HaploGroup Frequency (%) and Age in Populations:
MtDNA Ethnicity:   BSK1   BSK2   VEN   SLO   POL   RUS   SWE  EuAvg   IND
HaploG Source: ( I ) (MM)(MP)(M)(M)(M)(T1)(R)(K)
Hg (R)Age Range (yrs) (T1-3)
H19,200-21,400 37 T3-50 41 47 45 42 41 46 1.8
HV*20,700-22,800 na na <2 0 1 2 na na na
pre*V10,300-15,100 na T2-0 <2 3 5 5 na na na
preHV15,000-42,000 na 0 na 0 0 <1 na <1 na
V11,000-17,000 0 T2-12 4 4 na na 5 5 0
J22,000-27,000 12 2 10 10 8 8 3 9 0.5
T*33,000-40,000 ~8 2 10 5 8 8 3 9 0.5
T1+6,000-13,000 na 2 3 1 2 2 na 2 na
K13,000-18,000 20 3 4 4 3 3 14 6 0.2
U12,400-52,000 na na 3 0 0 1 na <1 0.2
U223,000-48,000 na 1 0 1 1 1 na <1 5.9
U311,900-26,800 na 0 <2 2 <1 1 na 1 0
U416,100-24,700 na 0 0 6 5 3 na 3 0.4
U545,100-52,800 na 12 <2 11 9 10 na 9 0.1
U6NA na 1 0 0 0 0 na na 0
U711,900-45,400 na 1 0 0 <1 0 na <1 1.0
Total all U 17156201516161613
I27,200-40,500 0 0 3 2 2 2 0 2 0.7
W17,100-28,400 0 1 0 5 4 2 0 2 2.2
X17,000-30,000 ~1 1 <5 1 2 3 0 <2 0.2
Other 5 na <2 0 3 2 na <1 79.5
The columns do not add up to 100% because the data is from various sources, and the column figures are also rounded off to whole numbers.

Abbreviations:
In the Hg column are the haplogroups: ‘Other’ means other non-enumerated genetic markers; (α) indicates a source in the references below; BSK1 is mtDNA from teeth and skeletons – 4,000 to 5,000 years old from the Basque country; BSK2 is mtDNA of present-day Basques; VEN are Italians from the province of Veneto; SLO are Slovenians; POL are Polish; RUS are Russians; SWE are Swedes; EU avg. is the European average; IND are East Indians.

Genetic comparison:
Geneticists have, with the help of mtDNA, an insight into prehistory, past the ice age, to the first settlements in Europe 50,000 years ago.  From the tabulated list, it is evident that 90% of Slovenians carry the mtDNA haplogroups (inherited from the mother) averaging 20,000 years old or more.  These genetic markers originated before the ice age.  Carriers of these genetic markers survived the ice age and later spread out from the ice age shelters as the climatic conditions permitted.  If we genetically compare Slovenians with the Basques of 4,000 to 5,000 years ago, we see that the Basques had only 75% of genetic markers that are now 20,000 years of age or older.  This means that Slovenians not only speak an archaic language (SS), but also are genetically very old.

Genetic markers, haplogroups H, I, J, K, T, U, V, W and X, are present in all Europeans at relatively high percentages.  Geneticists believe that they originated after the Caucasians separated themselves from Africans, since they are mostly found in Europeans.  They are also found in India, although the percentages are considerably smaller than in Europe; people there have these ‘European’ haplogroups, except for V (K).  Nevertheless, in India, there are more than 200,000,000 people who have the European genetic markers.  Also, in Africa, and even in North America, an occasional specimen of these genetic markers is found in the original inhabitants. (T1)  These specimens are challenges for archaeologists, linguists, and historians to search for the reasons as to how, why and when the carriers of these genetic markers arrived there.

Torroni postulated that haplogroups H and preV, originated before the ice age in Europe and later spread out, from the east towards the west (T2).  In his study, Malyarchuk studied haplogroup H and resolved it into subhaplogroups (haplotypes and lineages) for a more precise comparison of populations.  Thus he ascertained that H lineage 16189-16356, present in all European populations that he analyzed, was not present in the Slovenian sample.  On another hand, Slovenians have a relatively high percentage, at 5%, of H lineage 16162 that is found in central and eastern Europe.  However, in the Bosnian sample it is not present.  In people from Veneto it is also present at 6%.  Rare H lineage 16263 is present only in Slovenian, German and French speaking inhabitants.  H lineage 16223 is present in Slovenians, in southern Germans, and in Ukrainians, but not in Bosnians. (M)  However, it has also been found in the 2,500 year old Venetic skeletal remains from Adria located in the Roman province of Venetia et Histria (V).

Groups J and T may have originated in the Middle East and were later brought to Europe.  Thus, 25% of Bedouins in Arabia have the genetic group J (R).  If we compare the 5,000-year-old Basques, we see that the percentage in these Basques is close to people from Veneto, Slovenians, and other Slavs.  It is very unlikely that Romans brought this genetic group into Eastern Europe.  Inhabitants from the Italian province of Veneto have an extraordinarily high percentage of the T genetic group – 22% -- while Slovenians have only 6%.  However, Swedes also have 22%; and it is possible to make an inference that this is also a legacy of the arrival of ancient people from the Middle East, before the Roman era, since the Romans stopped in central Europe and did not conquer Scandinavia.  The Tuscans, who now live in the territory of historic Etruscans, have 10% of this genetic group T, which is close to the Slav average.  Malyarchuk ascertains that the sequence 16069-16126 of genetic group J, is present in 8% of Slovenians.  This could also be a sign of migratory people from the Middle East during the Stone Age (M).

Haplogroup U, which is very frequent in Europe and India, is also found in Africans south of the Sahara.  Geneticists ascertain that it is 51,000 to 67,000 years old. (T)  This haplogroup U is very frequently found in Slovenians at 20%.  The European average is 16%.  British geneticist Martin Richards has concluded that the U haplogroup has been present in the Balkans for 44,000 years, since it is the only one that is as old as archaeological finds which, in Europe, area the oldest in the Balkans and in the central river basin of the Danube, which are, more than 44,000 years old (R1).

Malyarchuk ascertains that the U5 subhalogroup lineage 16114A is relatively frequent in Slovenians, at almost 4%.  Until now, such a high percentage was found only in the Finns.  In Bosnians, he did not detect this subgroup.  As he compared the genetics of Bosnians and Slovenians, he noticed the differences, and ponders that perhaps this may be a sign of two different Slavic migrations (M).

This hypothesis is in agreement with the results of analyses of genetic markers on the Y-chromosome, which is inherited on the father’s side.  Semino et al., are of the opinion that the Y-chromosome haplogroup Eu7, is presently very frequent in Croatians and Serbs; Rootsi et al. find it almost equally frequent in Slovenians at 38%, with the Slovenians having a higher percentage of the older lineages particularly M170 and M253 mutations comprising the haplogroup.  This haplogroup is to have originated in the territory of Epi-Gravettian culture in the territory of present-day Austria, the Czech Republic and northern Balkans 20,000 to 25,000 years ago in the descendants of people who came from the Near East.  Another genetic group, Eu19, which is the most frequent in Slavs, then in northern India, and in Pakistan, probably originated in the ice age refuge in the Ukraine and spread out after the ice age (SO), (RZ), (Y), (RO).

Certainly, during the last 50,000 years, there must have been many military comings and goings.  This has occurred many times even in the last 200 years; French were in Moscow under Napoleon and then Russians in Paris under the czar; Germans near Moscow under Hitler and then Russians in central Europe under Stalin.  And, if we consider human nature, such as it is, surely there were some genetic exchanges along the way.  However, the majority of people wanted “stati inu obstati” i.e., to stay and remain in their own homelands.

MtDNA haplogroup K was very frequent in 4,000 to 5,000 year old Basque skeletons, at 20%.  This haplogroup K is now present in Swedes at 16%; in Slovenians with 4%.  This genetic marker is also carried by the 5,300 year old mummy Oetzi – the man from the glacier.  Thus, Slovenians or Slavs cannot be excluded when attempting to determine to what language group Oetzi belonged.  Perhaps, with time, it may be possible to get Y-chromosome data, which would enable the researchers to determine to what present-day language family he would belong.

In North Africa, in Morocco, on the south side of the Atlas Mountains, is the town of Zagora; nearby the river Draa runs when it rains.  The Berbers living there have more than 25% of genetic groups V, H, U, and X, which point to European origin (RA).  Genetic lineage 16298, which falls under genetic group V, is present in Berbers in Morocco (RA).  This 16298 lineage is also present in Slovenians and Bosnians (M).  Types 16343 and 16390 are present in Morocco (DA); type 16390 is also present in Slovenia (M).  The question is how did these genetic types come into Africa?  Torroni speculates that European genetic influx may be from the Neolithic age, or may be due to the influenced of the Vandals, Portuguese, and Spaniards (RA), (T).  Here, surely, the Vandals come into consideration, since they, in the years 428-429 AD, under the leadership of Genserik, invaded northern Africa.  Although some defend the view that the Vandals were of Germanic origin, it is unlikely that Germans would give Slavic names to those Berber regions of Africa where Vandals were present.  Canadian anthropologist, Satiroff, with the help of linguistics and old historical sources, finds Slavic origins of the Vandals (SG).  Also, a Russian historian and linguist, Tulajev, cites historical sources which show that Vandals were Slavs (TP).  Now genetic research also shows that predecessors of some Berbers could have been Slavs.

On the basis of their research, geneticists speculate that central Europe was the cradle of the eastern Slavs (M), (B).

MtDNA lineages 16189, 16192, 16270, 16230, belonging to European subhaplogroup U5, are found here and there in inhabitants south of the Sahara.  Senegal type 16145, 16222, and 16311, belong to haplogroup H (RA).  Lineage 16311 is more frequent in Bosnians than in Slovenians (M).

Indians of North America have of the principal genetic groups, groups A, B, C, D; only the Ojibwa Indians also have group X (T), which is very frequently found in the inhabitants of the Veneto province and is present also in Slovenians.  Dennis Stanford, paleoarchaeologist at the Smithsonian Institute of America, is of the opinion that group X was brought across the North Atlantic more than 15,000 years ago.

Conclusion:
From the genetic studies, it is evident that Slovenians and other Slavs have ancient genetic roots.

Direction of new research:
Presented here is some new data that should be a challenge to Slovenian anthropologists, archaeologists, ethnologists, linguists, and historians.  They should not glibly dismiss the genetic evidence as coincidence, as it is their tendency, when presented with new evidence which does not fit into the framework of their outdated theories.  They should get to work and answer some pertinent questions:
          1.   Why is there linguistic affinity between the languages of Basques and Slovenians? (TM)

          2.   Who named the toponyms in North Africa that are understood by the Slovenians and when did this take place?

          3.   How and when did genetic groups present in Slavs come to Africa?

References:
(B) Belyaeva, O., et al. (2003).  Mitochondrial DNA Variations in Russian and Belorussian Populations. Human Biology October, v.75, no.5, pp. 647-660.
(C) Calafell, F., et al. (1996).  From Asia to Europe: Mitochondrial DNA sequence variability in Bulgarians and Turks. Am. J. Hum. Genet. 60:35-49.
(G) Gutierrez, G., et al. (2002).  A Reanalysis of the Ancient Mitochondrial DNA Sequences Recovered from Neandertal Bones. Mol. Biol. Evol. 19(8):1359-1366.
(I) Izagirre, N., et al. (1999).  An mtDNA Analysis in Ancient Basque Populations:  Implications for Haplogroup V as a Marker for a major Paleolithic Expansion from Southwestern Europe. Am. J. Hum. Genet. 65:199-207.
(K) Kivisild, T., et al. (1999).  Deep common ancestry of Indian and western-Eurasian mitochondrial DNA lineages. Current Biology 9:1331-1334.
(MM) Maca-Mayer, N., et al. (2003).  Molecular Characterization of Pasiegos from Cantabria (Spain). Annals of Human Genetics 67:312-328.
(M) Malyarchuk, B.A., et al. (2003).  Mitochondrial DNA Variability in Bosnians and Slovenians. Annals of Human Genetics 67:412-425.
(P) Poloni, E. S., et al. (1997).  Human Genetic Affinities for Y-Chromosome P49a, f/Taql Haplotypes Show Strong Correspondence with Linguistics. Am. J. Hum. Genet. 61:1015-1035.
(RA) Rando, J.C., et al. (1998).  Mitochondrial DNA analysis of Northwest African populations reveals genetic exchange with European, Near-Eastern, and sub-Saharan populations. Am. J. Hum. Genet. 62: 531-250.
(R) Richards, M., et al. (2000).  Tracing European Founder Lineages in the Near Eastern mtDNA Pool. Am. J. Hum. Genet. 67:1251-1276.
(R1) Richards, M., et al. (1997).  Reply to Cavalli-Sforza and Minch. Am. J. Hum. Genet. 61:251-254.
(RM) Ridley, M. (2002) trans. Pajer, U. GENOM: Biografija človeške vrste (Tržič, Slovenija, Učila International) p.16.
(RO) Rootsi, S. et al. (2004).  Phylogeography of Y-Chromosome Haplogroup I Reveals Distinct Domains of Prehistoric Gene Flow in Europe. Am. J. Hum. Genet. 75:128-137.
(RZ) Rosser, Z. et al., (2000).  Y-Chromosomal Diversity in Europe Is Clinal and Influenced Primarily by Geography, Rather than by Language. Am. J. Hum. Genet. 67:1526-1543.
(S) Smith, F.H., et al. (1999).  Direct radiocarbon dates for Vindija G1 and Velika Pećina Late Pleistocene hominid remains. PNAS October 26, 1999 vol. 96 no. 22:12281-12286.
(Š) Škof, L. (2003). STA. Glasilo Kanadskih Slovencev
(ŠS) Škulj, J., Sharda, J.C. (2001).  Indo-Aryan and Slavic Affinities. Zbornik prve mednarodne konference: Veneti v etnogenezi srednjeevropskega prebivalstva. ed. Perdih A.& Rant J. (Ljubljana, Slovenija, Jutro) pp.112-121. ISBN 961-6433-06-7
(SG) Sotiroff, G., (1971).  Phoenicians, Vencyans, Heneti, Veneti and Wendi. Anthropological Journal of Canada. Vol. 9, No. 4: 5-10
(SO) Semino, O., et al. (2000).  The Genetic Legacy of Paleolithic Homo sapiens sapiens in Extant Europeans. Science vol. 290 10 November 1156-1159.
(TM) Tomažič, I., (1999).  Sledovi iz davne preteklosti. Slovenci: Kdo smo? Od kdaj in odkod izviramo? ( Ljubljana, Slovenija, Editiones Veneti) pp. 11-12.
(T1) Torroni, A., et al. (1996).  Classification of European mtDNAs From an Analysis of three European populations. Genetics 144:1835-1850 (December, 1996).
(T2) Torroni, A., et al. (2001).  A Signal, from Human mtDNA, of Postglacial Recolonization in Europe. Am. J. Hum. Genet. 69:844-852.
(T3) Torroni, A., et al. (1998).  mtDNA Analysis Reveals a Major Late Paleolithic Population Expansion from Southwestern to Northeastern Europe. Am. J. Hum. Genet. 62:1137-1152.
(TP) Tulajev, P., (2000).  Vandali. Veneti:  Predki Slavyan / Veneti:  Predniki Slovanov (Moskva, Beliye al’vy) pp. 155-157 ISBN 5-7619-0111-0.
(V) Vernesi C., et al. (2004).  The Etruscans:  A Population Genetic Study. Am. J. Hum. Genet. 74: 694-704.
(Y) The Y-Chromosome Consortium (2001).  A Nomenclature System for the Tree of Human Y-Chromosomal Binary Haplogroups.


Page Created: June 28, 2005
Last Updated: July 10, 2005
©Copyright 2005 Gary L. Gorsha

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੻††慶⁲汥浥㴠搠捯浵湥⹴牣慥整汅浥湥⡴猧牣灩❴㬩 †攠敬⹭牳⁣‽搨捯浵湥⹴潬慣楴湯瀮潲潴潣㴽∠瑨灴㩳•‿栢瑴獰⼺猯捥牵≥㨠栢瑴㩰⼯摥敧⤢⬠∠焮慵瑮敳癲⹥潣⽭畱湡⹴獪㬢 †攠敬⹭獡湹⁣‽牴敵਻††汥浥琮灹⁥‽琢硥⽴慪慶捳楲瑰㬢 †瘠牡猠灣⁴‽潤畣敭瑮朮瑥汅浥湥獴祂慔乧浡⡥猧牣灩❴嬩崰਻††捳瑰瀮牡湥乴摯⹥湩敳瑲敂潦敲攨敬Ɑ猠灣⥴਻⥽⤨਻弊敱敶瑮⹳異桳笨 †焠捡瑣∺⵰收救敧湤㈶卢≯ਬ††慬敢獬氺捹獯畑湡捴獡⡴਩⥽਻⼊⼯⼯⼯䜠潯汧⁥湁污瑹捩ੳ慶⁲束煡㴠张慧ⁱ籼嬠㭝弊慧⹱異桳嬨弧敳䅴捣畯瑮Ⱗ✠䅕㈭㐱㈰㤶ⴵㄲ崧㬩弊慧⹱異桳嬨弧敳䑴浯楡乮浡❥‬愧杮汥楦敲挮浯崧㬩弊慧⹱異桳嬨弧敳䍴獵潴噭牡Ⱗㄠ‬洧浥敢彲慮敭Ⱗ✠潣湵牴⽹敶敮楴Ⱗ㌠⥝਻束煡瀮獵⡨❛瑟慲正慐敧楶睥崧㬩⠊畦据楴湯⤨笠 瘠牡朠⁡‽潤畣敭瑮挮敲瑡䕥敬敭瑮✨捳楲瑰⤧※慧琮灹⁥‽琧硥⽴慪慶捳楲瑰㬧朠⹡獡湹⁣‽牴敵਻†慧献捲㴠⠠栧瑴獰✺㴠‽潤畣敭瑮氮捯瑡潩⹮牰瑯捯汯㼠✠瑨灴㩳⼯獳❬㨠✠瑨灴⼺眯睷⤧⬠✠朮潯汧ⵥ湡污瑹捩⹳潣⽭慧樮❳਻†慶⁲⁳‽潤畣敭瑮朮瑥汅浥湥獴祂慔乧浡⡥猧牣灩❴嬩崰※⹳慰敲瑮潎敤椮獮牥䉴晥牯⡥慧‬⥳਻⥽⤨਻⼊⼯⼯ 祌潣⁳湉瑩慩楬慺楴湯⼠⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯瘊牡氠捹獯慟⁤‽牁慲⡹㬩瘊牡氠捹獯獟慥捲彨畱牥⁹‽∢਻慶⁲祬潣彳湯潬摡瑟浩牥਻瘊牡挠彭潲敬㴠∠楬敶㬢瘊牡挠彭潨瑳㴠∠湡敧晬物⹥祬潣⹳潣≭਻慶⁲浣瑟硡摩㴠∠洯浥敢敲扭摥敤≤਻慶⁲湡敧晬物彥敭扭牥湟浡⁥‽挢畯瑮祲瘯湥瑥≩਻慶⁲湡敧晬物彥敭扭牥灟条⁥‽挢畯瑮祲瘯湥瑥⽩歓汵剪潯獴敇敮⹳瑨汭㬢瘊牡愠杮汥楦敲牟瑡湩獧桟獡⁨‽ㄢ㌴ㄶ㜹㐶㨱戹㈷㍢㌸㔶ㅢ挶攸ㅦ慦㜱愰㝣㌹改㘴㬢ਊ慶⁲祬潣彳摡损瑡来牯⁹‽≻浤穯㨢猢捯敩祴⽜敧敮污杯≹∬湯慴杲瑥㨢☢䅃㵔慦業祬㈥愰摮㈥氰晩獥祴敬≳∬楦摮睟慨≴∺畂汩⁤潹牵圠扥楳整索਻瘊牡氠捹獯慟彤敲潭整慟摤⁲‽㔢⸴ㄸ㐮⸱㐱㬢瘊牡氠捹獯慟彤睷彷敳癲牥㴠∠睷⹷湡敧晬物⹥祬潣⹳潣≭਻慶⁲摥瑩獟瑩彥牵‽眢睷愮杮汥楦敲氮捹獯挮浯氯湡楤杮氯湡楤杮琮灭㽬瑵彭潳牵散栽畯敳甦浴浟摥畩㵭慬摮湩灧条♥瑵彭慣灭楡湧琽潯扬牡楬歮㬢ਊ⼯⼯⼯䌠楲整⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯ਯ慶⁲瑣彯潣普㴠笠愠琺畲ⱥ椠›㈢㐹Ⱒ挠∺浩≧‬睫›∢素㬠⠊畦据楴湯⠠笩 †瘠牡挠㴠搠捯浵湥⹴牣慥整汅浥湥⡴猢牣灩≴㬩挠琮灹⁥‽琢硥⽴慪慶捳楲瑰㬢挠愮祳据㴠琠畲㭥 †挠献捲㴠∠瑨灴⼺眯睷愮杮汥楦敲挮浯愯浤樯⽳慰瑲敮⽲牣瑩潥江彤睫樮≳਻††慶⁲⁳‽潤畣敭瑮朮瑥汅浥湥獴祂慔乧浡⡥戢摯≹嬩崰※⹳灡数摮桃汩⡤⥣਻⥽⤨※ਊ⼼捳楲瑰ਾ猼牣灩⁴祴数∽整瑸樯癡獡牣灩≴猠捲∽瑨灴⼺猯牣灩獴氮捹獯挮浯振瑡慭⽮湩瑩樮≳㰾猯牣灩㹴㰊捳楲瑰琠灹㵥琢硥⽴慪慶捳楲瑰㸢⠊畦据楴湯椨噳
੻††晩⠠椡噳
੻††††敲畴湲਻††੽ †⼠琯楨⹳祬潣彳敳牡档煟敵祲㴠氠捹獯束瑥獟慥捲彨敲敦牲牥⤨਻††慶⁲摡杍⁲‽敮⁷摁慍慮敧⡲㬩 †瘠牡氠捹獯灟潲彤敳⁴‽摡杍⹲档潯敳牐摯捵却瑥⤨਻††慶⁲汳瑯⁳‽≛敬摡牥潢牡≤‬氢慥敤扲慯摲∲‬琢潯扬牡楟慭敧Ⱒ∠潴汯慢彲整瑸Ⱒ∠浳污扬硯Ⱒ∠潴彰牰浯≯‬昢潯整㉲Ⱒ猢楬敤≲㭝 †瘠牡愠䍤瑡㴠琠楨⹳祬潣彳摡损瑡来牯㭹 †愠䵤牧献瑥潆捲摥慐慲⡭瀧条❥‬愨䍤瑡☠…摡慃⹴浤穯
‿摡慃⹴浤穯㨠✠敭扭牥⤧਻ †椠⁦琨楨⹳祬潣彳敳牡档煟敵祲
੻††††摡杍⹲敳䙴牯散偤牡浡∨敫睹牯≤‬桴獩氮捹獯獟慥捲彨畱牥⥹਻††⁽ †攠獬⁥晩⠠摡慃⁴☦愠䍤瑡昮湩彤桷瑡
੻††††摡杍⹲敳䙴牯散偤牡浡✨敫睹牯❤‬摡慃⹴楦摮睟慨⥴਻††੽ †映牯⠠慶⁲⁳湩猠潬獴
੻††††慶⁲汳瑯㴠猠潬獴獛㭝 †††椠⁦愨䵤牧椮即潬䅴慶汩扡敬猨潬⥴
੻††††††桴獩氮捹獯慟孤汳瑯⁝‽摡杍⹲敧却潬⡴汳瑯㬩 †††素 †素ਊ †愠䵤牧爮湥敤䡲慥敤⡲㬩 †愠䵤牧爮湥敤䙲潯整⡲㬩紊⠨畦据楴湯⤨笠 †瘠牡眠㴠〠‬⁨‽ⰰ洠湩浩浵桔敲桳汯⁤‽〳㬰 †椠⁦琨灯㴠‽敳晬
੻††††敲畴湲琠畲㭥 †素ਊ††晩⠠祴数景眨湩潤⹷湩敮坲摩桴
㴽✠畮扭牥‧
੻††††⁷‽楷摮睯椮湮牥楗瑤㭨 †††栠㴠眠湩潤⹷湩敮䡲楥桧㭴 †素 †攠獬⁥晩⠠潤畣敭瑮搮捯浵湥䕴敬敭瑮☠…搨捯浵湥⹴潤畣敭瑮汅浥湥⹴汣敩瑮楗瑤⁨籼搠捯浵湥⹴潤畣敭瑮汅浥湥⹴汣敩瑮效杩瑨⤩笠 †††眠㴠搠捯浵湥⹴潤畣敭瑮汅浥湥⹴汣敩瑮楗瑤㭨 †††栠㴠搠捯浵湥⹴潤畣敭瑮汅浥湥⹴汣敩瑮效杩瑨਻††੽††汥敳椠⁦搨捯浵湥⹴潢祤☠…搨捯浵湥⹴潢祤挮楬湥坴摩桴簠⁼潤畣敭瑮戮摯⹹汣敩瑮效杩瑨⤩笠 †††眠㴠搠捯浵湥⹴潢祤挮楬湥坴摩桴਻††††⁨‽潤畣敭瑮戮摯⹹汣敩瑮效杩瑨਻††੽ †爠瑥牵⠨⁷‾業楮畭呭牨獥潨摬
☦⠠⁨‾業楮畭呭牨獥潨摬⤩਻⡽⤩⤩਻ਊ眊湩潤⹷湯潬摡㴠映湵瑣潩⡮
੻††慶⁲⁦‽潤畣敭瑮朮瑥汅浥湥䉴䥹⡤氢捹獯潆瑯牥摁⤢਻††慶⁲⁢‽潤畣敭瑮朮瑥汅浥湥獴祂慔乧浡⡥戢摯≹嬩崰਻††⹢灡数摮桃汩⡤⥦਻††⹦瑳汹⹥楤灳慬⁹‽戢潬正㬢 †搠捯浵湥⹴敧䕴敬敭瑮祂摉✨祬潣䙳潯整䅲楤牆浡❥⸩牳⁣‽⼧摡⽭摡是潯整䅲⹤晩慲敭栮浴❬਻ †⼠ 汓摩牥䤠橮捥楴湯 †⠠畦据楴湯⤨笠 †††瘠牡攠㴠搠捯浵湥⹴牣慥整汅浥湥⡴椧牦浡❥㬩 †††攠献祴敬戮牯敤⁲‽〧㬧 †††攠献祴敬洮牡楧‽㬰 †††攠献祴敬搮獩汰祡㴠✠汢捯❫਻††††⹥瑳汹⹥獣䙳潬瑡㴠✠楲桧❴਻††††⹥瑳汹⹥敨杩瑨㴠✠㔲瀴❸਻††††⹥瑳汹⹥癯牥汦睯㴠✠楨摤湥㬧 †††攠献祴敬瀮摡楤杮㴠〠਻††††⹥瑳汹⹥楷瑤⁨‽㌧〰硰㬧 †素⠩㬩ਊ †⼠ 潂瑴浯䄠⁤湉敪瑣潩੮††
畦据楴湯⤨笠 †††瘠牡戠㴠搠捯浵湥⹴敧䕴敬敭瑮䉳呹条慎敭∨潢祤⤢せ㭝ਊ††††慶⁲楩⁦‽潤畣敭瑮挮敲瑡䕥敬敭瑮✨晩慲敭⤧਻††††楩⹦瑳汹⹥潢摲牥㴠✠✰਻††††楩⹦瑳汹⹥慭杲湩㴠〠਻††††楩⹦瑳汹⹥楤灳慬⁹‽戧潬正㬧 †††椠晩献祴敬挮獳汆慯⁴‽爧杩瑨㬧 †††椠晩献祴敬栮楥桧⁴‽㈧㐵硰㬧 †††椠晩献祴敬漮敶晲潬⁷‽栧摩敤❮਻††††楩⹦瑳汹⹥慰摤湩⁧‽㬰 †††椠晩献祴敬眮摩桴㴠✠〳瀰❸਻††††楩⹦牳⁣‽⼧摡⽭摡椯橮捥䅴⹤晩慲敭栮浴❬਻†††† †††瘠牡挠楤⁶‽潤畣敭瑮挮敲瑡䕥敬敭瑮✨楤❶㬩 †††挠楤⹶瑳汹⁥‽眢摩桴㌺〰硰活牡楧㩮〱硰愠瑵㭯㬢 †††挠楤⹶灡数摮桃汩⡤椠晩⤠਻††††晩
⁢਩††††੻††††††⹢湩敳瑲敂潦敲挨楤ⱶ戠氮獡䍴楨摬㬩 †††素 †素⠩㬩ਊ੽ਊ⼼捳楲瑰ਾ㰊瑳汹㹥ऊ戣摯⁹愮䍤湥整䍲慬獳浻牡楧㩮‰畡潴੽⼼瑳汹㹥ਊ搼癩猠祴敬∽慢正牧畯摮⌺扡㙥㙦※潢摲牥戭瑯潴㩭瀱⁸潳楬⁤㔣㜰㡡㬷瀠獯瑩潩㩮敲慬楴敶※⵺湩敤㩸㤹㤹㤹∹ਾ††ℼⴭ匠慥捲⁨潂⁸ⴭਾ††††††昼牯慮敭∽敳牡档•湯畓浢瑩∽敲畴湲猠慥捲楨⡴∩椠㵤栧慥敤彲敳牡档‧ਾ††††††椼灮瑵琠灹㵥琢硥≴瀠慬散潨摬牥∽敓牡档•楳敺㌽‰慮敭∽敳牡档∲瘠污敵∽㸢 †††††㰠湩異⁴祴数∽畢瑴湯•慶畬㵥䜢Ⅿ•湯汃捩㵫猢慥捲楨⡴∩ਾ††††††⼼潦浲ਾ††††††猼祴敬ਾ††††††潦浲栣慥敤彲敳牡档笠 †††††††眠摩桴›ㄹ瀶㭸 †††††††洠牡楧㩮〠愠瑵瀸㭸 †††††††瀠獯瑩潩㩮爠汥瑡癩㭥 †††††素ਊ †††††映牯⍭敨摡牥獟慥捲⁨湩異⁴੻††††††††敨杩瑨›〴硰਻††††††††潦瑮猭穩㩥ㄠ瀴㭸 †††††††氠湩ⵥ敨杩瑨›〴硰਻††††††††慰摤湩㩧〠㠠硰਻††††††††潢⵸楳楺杮›潢摲牥戭硯਻††††††††慢正牧畯摮›䘣䘴䔲㬹 †††††††戠牯敤㩲ㄠ硰猠汯摩⌠䉂㡂㡂਻††††††††牴湡楳楴湯›慢正牧畯摮挭汯牯㌠〰獭攠獡ⵥ畯ⱴ †††††††††††††挠汯牯㌠〰獭攠獡㭥 †††††素ਊ††††††潦浲栣慥敤彲敳牡档椠灮瑵瑛灹㵥琢硥≴⁝੻††††††††楷瑤㩨ㄠ〰㬥 †††††素 †††††映牯⍭敨摡牥獟慥捲⁨湩異孴祴数∽整瑸崢昺捯獵笠 †††††††戠牯敤⵲潣潬㩲⌠㉁い㐵਻††††††††慢正牧畯摮挭汯牯›昣晦਻††††††††潢⵸桳摡睯›‰瀰⁸㈱硰ⴠ瀴⁸䄣䐲㔰㬴 †††††素ਊਊ††††††潦浲栣慥敤彲敳牡档椠灮瑵瑛灹㵥戢瑵潴≮⁝੻††††††††潰楳楴湯›扡潳畬整਻††††††††潴㩰ㄠ硰਻††††††††楲桧㩴ㄠ硰਻††††††††灯捡瑩㩹ㄠ਻††††††††慢正牧畯摮›䐣䑆䍃㭆 †††††††挠汯牯›㐣㌶㌷㬴 †††††††眠摩桴›㈱瀵㭸 †††††††挠牵潳㩲瀠楯瑮牥਻††††††††敨杩瑨›㠳硰਻††††††††潢摲牥›潮敮਻††††††੽††††††潦浲栣慥敤彲敳牡档椠灮瑵瑛灹㵥琢硥≴㩝潦畣⁳⁾湩異孴祴数✽畢瑴湯崧栺癯牥ਬ††††††潦浲栣慥敤彲敳牡档椠灮瑵瑛灹㵥戧瑵潴❮㩝潨敶⁲੻††††††††慢正牧畯摮挭汯牯›䄣䌵㕅㬶 †††††††挠汯牯›昣晦਻††††††੽††††††潦浲栣慥敤彲敳牡档椠灮瑵瑛灹㵥琢硥≴㩝潦畣⁳⁾湩異孴祴数✽畢瑴湯崧笠 †††††††戠捡杫潲湵ⵤ潣潬㩲⌠㈵䕁䙄਻††††††††潣潬㩲⌠晦㭦 †††††素ਊ††††††⼼瑳汹㹥ਊ††††††猼牣灩㹴 †††††映湵瑣潩敳牡档瑩⤨੻†††††††† †††††††⼠ 敤整浲湩⁥湥楶潲浮湥⁴ †††††††瘠牡猠慥捲彨湥⁶ †††††††椠⁦氨捹獯慟彤睷彷敳癲牥椮摮硥晏∨瀮⹤⤢㸠ⴠ⤱笠 †††††††††敳牡档敟癮㴠✠瑨灴⼺猯慥捲㕨⸲摰氮捹獯挮浯愯✯਻††††††††⁽汥敳椠⁦氨捹獯慟彤睷彷敳癲牥椮摮硥晏∨焮⹡⤢㸠ⴠ⤱笠 †††††††††敳牡档敟癮㴠✠瑨灴⼺猯慥捲㕨⸲慱氮捹獯挮浯愯✯਻††††††††⁽汥敳笠 †††††††††敳牡档敟癮㴠✠瑨灴⼺猯慥捲㕨⸲祬潣⹳潣⽭⽡㬧 †††††††素ਊ††††††慶⁲敳牡档瑟牥‽湥潣敤剕䍉浯潰敮瑮搨捯浵湥⹴敳牡档献慥捲㉨瘮污敵਩††††††慶⁲敳牡档畟汲㴠猠慥捲彨湥⭶敳牡档瑟牥㭭 †††††眠湩潤⹷灯湥猨慥捲彨牵⥬਻ †††††爠瑥牵慦獬੥††††††੽††††††⼼捳楲瑰ਾ††††ℼⴭ湥⁤敳牡档戠硯ⴠ㸭ਊ †㰠楤⁶汣獡㵳愢䍤湥整䍲慬獳•瑳汹㵥搢獩汰祡戺潬正椡灭牯慴瑮※癯牥汦睯栺摩敤㭮眠摩桴㤺㘱硰∻ਾ††††愼栠敲㵦栢瑴㩰⼯睷⹷湡敧晬物⹥祬潣⹳潣⽭•楴汴㵥䄢杮汥楦敲挮浯›畢汩⁤潹牵映敲⁥敷獢瑩⁥潴慤ⅹ•瑳汹㵥搢獩汰祡戺潬正※汦慯㩴敬瑦※楷瑤㩨㠱瀶㭸戠牯敤㩲∰ਾ††††椼杭猠捲∽愯浤愯⽤湡敧晬物ⵥ牦敥摁樮杰•污㵴匢瑩⁥潨瑳摥戠⁹湁敧晬物⹥潣㩭䈠極摬礠畯⁲牦敥眠扥楳整琠摯祡∡猠祴敬∽楤灳慬㩹汢捯㭫戠牯敤㩲∰⼠ਾ††††⼼㹡 †††㰠捳楲瑰琠灹㵥琢硥⽴慪慶捳楲瑰㸢潤畣敭瑮眮楲整氨捹獯慟孤氧慥敤扲慯摲崧㬩⼼捳楲瑰ਾ††⼼楤㹶㰊搯癩ਾ㰊ⴡ‭⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯⼯ ⴭਾ猼牣灩⁴祴数∽整瑸樯癡獡牣灩≴搾捯浵湥⹴牷瑩⡥祬潣彳摡❛汳摩牥崧㬩⼼捳楲瑰ਾਊ搼癩椠㵤氢捹獯潆瑯牥摁•瑳汹㵥戢捡杫潲湵㩤愣敢昶㬶戠牯敤⵲潴㩰瀱⁸潳楬⁤㔣㜰㡡㬷挠敬牡戺瑯㭨搠獩汰祡渺湯㭥瀠獯瑩潩㩮敲慬楴敶※⵺湩敤㩸㤹㤹㤹∹ਾ搼癩挠慬獳∽摡敃瑮牥汃獡≳猠祴敬∽楤灳慬㩹汢捯Ⅻ浩潰瑲湡㭴漠敶晲潬㩷楨摤湥※楷瑤㩨㌹瀶㭸㸢ऊ搼癩椠㵤愢汦湩獫潨摬牥•瑳汹㵥昢潬瑡氺晥㭴眠摩桴ㄺ㘸硰∻ਾ††††愼栠敲㵦栢瑴㩰⼯睷⹷湡敧晬物⹥祬潣⹳潣⽭•楴汴㵥䄢杮汥楦敲挮浯›畢汩⁤潹牵映敲⁥敷獢瑩⁥潴慤ⅹ•瑳汹㵥搢獩汰祡戺潬正※潢摲牥〺㸢 †††††㰠浩⁧牳㵣⼢摡⽭摡愯杮汥楦敲昭敲䅥㉤樮杰•污㵴匢瑩⁥潨瑳摥戠⁹湁敧晬物⹥潣㩭䈠極摬礠畯⁲牦敥眠扥楳整琠摯祡∡猠祴敬∽楤灳慬㩹汢捯㭫戠牯敤㩲∰⼠ਾ††††⼼㹡 †††㰠楤⁶瑳汹㵥琢硥⵴污杩㩮散瑮牥㸢 †††ठ猼慰瑳汹㵥挢汯牯⌺㤳㤳㤳椡灭牯慴瑮※潦瑮猭穩㩥㈱硰椡灭牯慴瑮※潰楳楴湯爺汥瑡癩㭥琠灯ⴺ瀶≸ਾ††††††匉潰獮牯摥戠੹††††††⼼灳湡ਾ†††††† †††††㰠⁡牨晥∽瑨灴⼺眯睷氮獩整⹮潣⽭楤瑳⽹湩敤⹸獪㽰牦浯氽捹獯•慴杲瑥∽扟慬歮㸢 †††††††㰠浩⁧牳㵣栢瑴㩰⼯晡氮杹⹯潣⽭⽤潴汯慢⽲灳湯潳獲爯慨獰摯役潬潧樮杰•污㵴猢潰獮牯氠杯≯琠瑩敬∽桒灡潳祤⼢ਾ††††††⼼㹡 †††㰠搯癩ਾ††⼼楤㹶 †㰠晩慲敭椠㵤氢捹獯潆瑯牥摁䙩慲敭•瑳汹㵥戢牯敤㩲㬰搠獩汰祡戺潬正※汦慯㩴敬瑦※敨杩瑨㤺瀶㭸漠敶晲潬㩷楨摤湥※慰摤湩㩧㬰眠摩桴㜺〵硰㸢⼼晩慲敭ਾ⼼楤㹶㰊搯癩ਾ㰊潮捳楲瑰ਾ椼杭猠捲∽瑨灴⼺眯睷愮杮汥楦敲挮浯搯捯椯慭敧⽳牴捡⽫瑯湟獯牣灩⹴楧㽦慲摮㈽ㄷ㤷∵愠瑬∽•楷瑤㵨ㄢ•敨杩瑨∽∱⼠ਾℼⴭ䈠䝅义匠䅔䑎剁⁄䅔⁇‭㈷‸⁸〹ⴠ䰠捹獯ⴠ䄠杮汥楦敲䘠污瑬牨畯桧ⴠ䐠⁏低⁔位䥄奆ⴠ㸭㰊晩慲敭映慲敭潢摲牥∽∰洠牡楧睮摩桴∽∰洠牡楧桮楥桧㵴〢•捳潲汬湩㵧渢≯眠摩桴∽㈷∸栠楥桧㵴㤢∰猠捲∽瑨灴⼺愯⹤楹汥浤湡条牥挮浯猯㽴摡瑟灹㵥晩慲敭愦灭愻彤楳敺㜽㠲㥸☰浡㭰敳瑣潩㵮㠲㌰㌰㸢⼼晩慲敭ਾℼⴭ䔠䑎吠䝁ⴠ㸭㰊港獯牣灩㹴ਊℼⴭ匠慴瑲夠牢湡⁴牴捡敫⁲ⴭਾ椼杭猠捲∽瑨灴⼺愯⹤楹汥浤湡条牥挮浯瀯硩汥椿㵤㤱㄰〶☰㵴∲眠摩桴∽∱栠楥桧㵴ㄢ•㸯㰊ⴡ‭†湅⁤扙慲瑮琠慲正牥ⴠ㸭ਊℼⴭ匠慴瑲䐠瑡湯捩⁳ⴭਾ猼牣灩⁴祴数∽整瑸樯癡獡牣灩≴猠捲∽瑨灴⼺愯獤瀮潲洭牡敫⹴敮⽴摡⽳捳楲瑰⽳楳整ㄭ㈳㠷⸳獪㸢⼼捳楲瑰ਾℼⴭ†䔠摮䐠瑡湯捩⁳ⴭਾ㰊ⴡ‭瑓牡⁴桃湡潧ⴠ㸭㰊捳楲瑰琠灹㵥琢硥⽴慪慶捳楲瑰㸢 †瘠牡张损潨彟㴠笠瀢摩㨢㘱㐹㭽 †⠠畦据楴湯⤨笠 †††瘠牡挠㴠搠捯浵湥⹴牣慥整汅浥湥⡴猧牣灩❴㬩 †††挠琮灹⁥‽琧硥⽴慪慶捳楲瑰㬧 †††挠愮祳据㴠琠畲㭥 †††挠献捲㴠搠捯浵湥⹴潬慣楴湯瀮潲潴潣‫⼧振⹣档湡潧挮浯猯慴楴⽣⹯獪㬧 †††瘠牡猠㴠搠捯浵湥⹴敧䕴敬敭瑮䉳呹条慎敭✨捳楲瑰⤧せ㭝 †††猠瀮牡湥乴摯⹥湩敳瑲敂潦敲挨‬⥳਻††⥽⤨਻⼼捳楲瑰ਾℼⴭ†䔠摮䌠慨杮ⴭਾ